Base of the evolutionary tree for Chactoidea

← parent

• Chactoidea (531 species)
  • Chactidae → (206 species)
    • Brotheinae → (34 species)
      • Brotheini → (29 species)
        • Brotheas humboldti
        • Brotheas bolivianus
        • Brotheas ocamoi
        • Brotheas caramaschii
        • Brotheas libinallyi
        • Brotheas munozi
        • Brotheas mingueti
        • Brotheas granimanus
        • Brotheas camposi
        • Brotheas wareipai
        • Brotheas cataniapensis
        • Brotheas wilmeri
        • Brotheas cristinae
        • Brotheas silvestris
        • Brotheas gervaisii
        • Brotheas lichyi
        • Brotheas overali
        • Brotheas granulatus
        • Brotheas sanabriai
        • Brotheas rionegroensis
        • Brotheas cunucunumensis
        • Brotheas paraensis
        • Brotheas henriquesi
        • Brotheas dasilvai
        • Brotheas mawarinumensis
        • Brotheas amazonicus
        • Brotheas jourdani
        • Brotheas perezramirezi
        • Brotheas tapajos
      • Belisariini → (5 species)
        • Troglotayosicus → (3 species)
        • Belisarius → (2 species)
    • Teuthraustes → (23 species)
      • Teuthraustes rosenbergi
      • Teuthraustes camposi
      • Teuthraustes lojanus
      • Teuthraustes adrianae
      • Teuthraustes atramentarius
      • Teuthraustes amazonicus
      • Teuthraustes reticulatus
      • Teuthraustes braziliensis
      • Teuthraustes gervaisii
      • Teuthraustes carmelinae
      • Teuthraustes guerdouxi
      • Teuthraustes dubius
      • Teuthraustes oculatus
      • Teuthraustes whymperi
      • Teuthraustes ohausi
      • Teuthraustes maturaca
      • Teuthraustes lisei
      • Teuthraustes akananensis
      • Teuthraustes simonsi
      • Teuthraustes newaribe
      • Teuthraustes festae
      • Teuthraustes wittii
      • Teuthraustes glaber
    • Chactopsoides → (4 species)
      • Chactopsoides marahuacaensis
      • Chactopsoides anduzei
      • Chactopsoides gonzalezspongai
      • Chactopsoides yanomami
    • Vachoniochactas → (5 species)
      • Vachoniochactas humboldti
      • Vachoniochactas amazonicus
      • Vachoniochactas ashleeae
      • Vachoniochactas roraima
      • Vachoniochactas lasallei
    • Chactinae → (52 species)
      • Chactas latuffi
      • Chactas chrysopus
      • Chactas oxfordi
      • Chactas venegasi
      • Chactas platillonensis
      • Chactas lepturus
      • Chactas gansi
      • Chactas interpuncta
      • Chactas barbacoensis
      • Chactas moreti
      • Chactas simonii
      • Chactas mahnerti
      • Chactas reticulatus
      • Chactas koepckei
      • Chactas brevicaudatus
      • Chactas rogelioi
      • Chactas exsul
      • Chactas gestroi
      • Chactas choroniensis
      • Chactas mauriesi
      • Chactas chabasquensis
      • Chactas braziliensis
      • Chactas barravierai
      • Chactas raymondhansi
      • Chactas keyserlingii
      • Chactas rubrolineatus
      • Chactas viloriai
      • Chactas yaupi
      • Chactas maimirensis
      • Chactas marinae
      • Chactas ferruginosus
      • Chactas setosus
      • Chactas aequinoctialis
      • Chactas adornellae
      • Chactas iutensis
      • Chactas yupai
      • Chactas vanbenedenii
      • Chactas bonito
      • Chactas brownelli
      • Chactas campoelisasensis
      • Chactas major
      • Chactas scabrimanus
      • Chactas ozendai
      • Chactas alarconi
      • Chactas guinandcortesi
      • Chactas tumaquensis
      • Chactas karschii
      • Chactas laevipes
      • Chactas hatilloensis
      • Chactas turguaensis
      • Chactas granulosus
      • Chactas hauseri
    • Neochactas → (38 species)
      • Neochactas sanmartini
      • Neochactas bilbaoi
      • Neochactas yekuanae
      • Neochactas bariensis
      • Neochactas riopinensis
      • Neochactas paoensis
      • Neochactas gaillardi
      • Neochactas mottai
      • Neochactas caroniensis
      • Neochactas santanai
      • Neochactas leoneli
      • Neochactas garciai
      • Neochactas eliasilvai
      • Neochactas laui
      • Neochactas jaspei
      • Neochactas fei
      • Neochactas parvulus
      • Neochactas sarisarinamensis
      • Neochactas panarei
      • Neochactas guaiquinimensis
      • Neochactas granosus
      • Neochactas neblinensis
      • Neochactas macrochelae
      • Neochactas ruizpittoli
      • Neochactas kjellesvigi
      • Neochactas simarawochensis
      • Neochactas verai
      • Neochactas bruzuali
      • Neochactas racenisi
      • Neochactas delicatus
      • Neochactas kelleri
      • Neochactas colombiensis
      • Neochactas sissomi
      • Neochactas fravalae
      • Neochactas josemanueli
      • Neochactas skuki
      • Neochactas efreni
      • Neochactas orinocensis
    • Uroctoninae → (5 species)
      • Uroctonus → (3 species)
        • Uroctonus franckei
        • western forest scorpion (Uroctonus mordax)
        • Uroctonus grahami
      • Anuroctonus → (2 species)
        • Anuroctonus phaiodactylus
        • Anuroctonus pococki
    • Broteochactas → (15 species)
      • Broteochactas venezuelensis
      • Broteochactas trezzii
      • Broteochactas manisapanensis
      • Broteochactas gollmeri
      • Broteochactas parimensis
      • Broteochactas verneti
      • Broteochactas danielleae
      • Broteochactas nitidus
      • Broteochactas silves
      • Broteochactas porosus
      • Broteochactas cocuyensis
      • Broteochactas cauaburi
      • Broteochactas niemeyerae
      • Broteochactas scorzai
      • Broteochactas vestigialis
    • Hadrurochactas → (8 species)
      • Hadrurochactas araripe
      • Hadrurochactas odoardoi
      • Hadrurochactas machadoi
      • Hadrurochactas cristinae
      • Hadrurochactas polisi
      • Hadrurochactas schaumii
      • Hadrurochactas brejo
      • Hadrurochactas mapuera
    • Chactopsis → (9 species)
      • Chactopsis carolinae
      • Chactopsis barajuri
      • Chactopsis curupira
      • Chactopsis siapaensis
      • Chactopsis insignis
      • Chactopsis amazonica
      • Chactopsis chullachaqui
      • Chactopsis sujirima
      • Chactopsis buhrnheimi
    • Megachactops → (2 species)
      • Megachactops coriacea
      • Megachactops kuemoi
    • Auyantepuia → (5 species)
      • Auyantepuia amapaensis
      • Auyantepuia laurae
      • Auyantepuia aluku
      • Auyantepuia aurum
      • Auyantepuia surinamensis
    • Guyanochactas → (5 species)
      • Guyanochactas gonzalezspongai
      • Guyanochactas flavus
      • Guyanochactas goujei
      • Guyanochactas mascarenhasi
      • Guyanochactas touroulti
    • Spinochactas mitaraka
  • Vaejovidae → (196 species)
    • Vaejovis → (73 species)
      • Vaejovis tenuipalpus
      • Vaejovis gracilis
      • Vaejovis curvidigitus
      • Vaejovis paysonensis
      • Vaejovis tapalpa
      • Vaejovis cashi
      • Vaejovis talpa
      • Vaejovis zapoteca
      • Vaejovis deboerae
      • Vaejovis chiapas
      • Vaejovis montanus
      • Vaejovis punctatus
      • Vaejovis dugesi
      • Vaejovis morelia
      • Vaejovis prendinii
      • Vaejovis monticola
      • Vaejovis franckei
      • Vaejovis bruneus
      • Vaejovis electrum
      • Vaejovis bigelowi
      • Vaejovis glabrimanus
      • Vaejovis darwini
      • Vaejovis vorhiesi
      • Vaejovis feti
      • Vaejovis santibagnezi
      • Vaejovis decipiens
      • Vaejovis coalcoman
      • Vaejovis granulatus
      • Vaejovis mauryi
      • Vaejovis sprousei
      • Vaejovis davidi
      • Vaejovis oaxaca
      • Vaejovis pusillus
      • Vaejovis intrepidus
      • Vaejovis yosemitensis
      • Vaejovis solegladi
      • Vaejovis nigrescens
      • Vaejovis trinityae
      • Southern Unstriped Scorpion (Vaejovis carolinianus)
      • Vaejovis grayae
      • Vaejovis nigrofemoratus
      • Vaejovis smithi
      • Vaejovis brysoni
      • Vaejovis ocotensis
      • Vaejovis mcwesti
      • Vaejovis dzahui
      • Vaejovis oaxacae
      • Vaejovis pequeno
      • Vaejovis ceboruco
      • Vaejovis janssi
      • Vaejovis vaquero
      • Vaejovis maculosus
      • Vaejovis intermedius
      • Vaejovis chisos
      • Vaejovis setosus
      • Vaejovis rossmani
      • Vaejovis mitchelli
      • Vaejovis troupi
      • Vaejovis bandido
      • Vaejovis lapidicola
      • Vaejovis nanchititla
      • Vaejovis sierrae
      • Vaejovis trespicos
      • Vaejovis mexicanus
      • Vaejovis jonesi
      • Vaejovis crumpi
      • Vaejovis norteno
      • Vaejovis tesselatus
      • Vaejovis tenamaztlei
      • Vaejovis grahami
      • Vaejovis halli
      • Vaejovis flavus
      • Vaejovis pococki
    • Pseudouroctonus → (13 species)
      • Pseudouroctonus glimmei
      • Pseudouroctonus peccatum
      • Pseudouroctonus minimus
      • Pseudouroctonus reddelli
      • Pseudouroctonus andreas
      • Pseudouroctonus chicano
      • Pseudouroctonus saavasi
      • Pseudouroctonus sprousei
      • Pseudouroctonus cazieri
      • Pseudouroctonus lindsayi
      • Pseudouroctonus iviei
      • Pseudouroctonus apacheanus
      • Pseudouroctonus rufulus
    • Kochius → (9 species)
      • Kochius villosus
      • Kochius sonorae
      • Kochius barbatus
      • Kochius magdalensis
      • Kochius cerralvensis
      • Kochius punctipalpi
      • Kochius bruneus
      • Kochius hirsuticauda
      • Kochius insularis
    • Syntropis → (4 species)
      • Syntropis macrura
      • Syntropis longiunguis
      • Syntropis williamsi
      • Syntropis aalbui
    • Gertschius → (2 species)
      • Gertschius agilis
      • Gertschius crassicorpus
    • Franckeus → (6 species)
      • Franckeus kochi
      • Franckeus peninsularis
      • Franckeus nitidulus
      • Franckeus rubrimanus
      • Franckeus minckleyi
      • Franckeus platnicki
    • Paravaejovis → (11 species)
      • Paravaejovis confusus
      • Paravaejovis schwenkmeyeri
      • Paravaejovis waeringi
      • Paravaejovis gravicaudus
      • Paravaejovis pumilis
      • Paravaejovis puritanus
      • Paravaejovis diazi
      • Paravaejovis eusthenura
      • Paravaejovis hoffmanni
      • Paravaejovis spinigerus
      • Paravaejovis galbus
    • Thorellius → (3 species)
      • Thorellius atrox
      • Thorellius intrepidus
      • Thorellius cristimanus
    • Stahnkeus → (5 species)
      • Stahnkeus deserticola
      • Stahnkeus harbisoni
      • Stahnkeus subtilimanus
      • Stahnkeus allredi
      • Stahnkeus polisi
    • Uroctonites → (4 species)
      • Uroctonites montereus
      • Uroctonites sequoia
      • Uroctonites giulianii
      • Uroctonites huachuca
    • Paruroctonus → (30 species)
      • Paruroctonus utahensis
      • Paruroctonus marksi
      • Paruroctonus coahuilanus
      • Paruroctonus arnaudi
      • Paruroctonus bajae
      • Northern Scorpion (Paruroctonus boreus)
      • Paruroctonus xanthus
      • Monterey Dunes Scorpion (Paruroctonus maritimus)
      • Paruroctonus pseudopumilis
      • Paruroctonus bantai
      • Paruroctonus ventosus
      • Paruroctonus hirsutipes
      • Paruroctonus nitidus
      • Paruroctonus williamsi
      • Paruroctonus shulovi
      • Paruroctonus variabilis
      • Paruroctonus boquillas
      • Paruroctonus pecos
      • Paruroctonus stahnkei
      • Paruroctonus borregoensis
      • Paruroctonus ammonastes
      • Paruroctonus becki
      • Paruroctonus simulatus
      • Paruroctonus surensis
      • Paruroctonus baergi
      • Paruroctonus silvestrii
      • Paruroctonus gracilior
      • Paruroctonus arenicola
      • Paruroctonus mesaensis
      • Paruroctonus luteolus
    • Smeringurus → (4 species)
      • Smeringurus mesaensis
      • Smeringurus grandis
      • Smeringurus vachoni
      • Smeringurus aridus
    • Wernerius → (3 species)
      • Wernerius inyoensis
      • Wernerius spicatus
      • Wernerius mumai
    • Mesomexovis → (7 species)
      • Mesomexovis subcristatus
      • Mesomexovis punctatus
      • Mesomexovis atenango
      • Mesomexovis occidentalis
      • Mesomexovis spadix
      • Mesomexovis variegatus
      • Mesomexovis oaxaca
    • Chihuahuanus → (8 species)
      • Chihuahuanus kovariki
      • Chihuahuanus glabrimanus
      • Chihuahuanus crassimanus
      • Chihuahuanus cazieri
      • Chihuahuanus russelli
      • Chihuahuanus globosus
      • Chihuahuanus bilineatus
      • Chihuahuanus coahuilae
    • Konetontli → (5 species)
      • Konetontli pattersoni
      • Konetontli acapulco
      • Konetontli nayarit
      • Konetontli chamelaensis
      • Konetontli kuarapu
    • Kovarikia → (4 species)
      • Kovarikia oxy
      • Kovarikia angelena
      • Kovarikia bogerti
      • Kovarikia savaryi
    • Maaykuyak → (2 species)
      • Maaykuyak vittatus
      • Maaykuyak waueri
    • Balsateres cisnerosi
    • Kuarapu purhepecha
    • Vizcaino viscainensis
  • Euscorpiidae → (128 species)
    • Scorpiopinae → (56 species)
      • Scorpiops → (29 species)
        • Scorpiops ingens
        • Scorpiops rohtangensis
        • Scorpiops leptochirus
        • Scorpiops pachmarhicus
        • Scorpiops afghanus
        • Scorpiops lhasa
        • Scorpiops pakistanus
        • Scorpiops kraepelini
        • Scorpiops atomatus
        • Scorpiops zubairahmedi
        • Scorpiops pseudomontanus
        • Scorpiops oligotrichus
        • Scorpiops luridus
        • Scorpiops dentidactylus
        • Scorpiops braunwalderi
        • Scorpiops spitiensis
        • Scorpiops feti
        • Scorpiops lindbergi
        • Scorpiops hardwickii
        • Scorpiops demisi
        • Scorpiops dastychi
        • Scorpiops jendeki
        • Scorpiops pococki
        • Scorpiops irenae
        • Scorpiops langxian
        • Scorpiops petersii
        • Scorpiops margerisonae
        • Scorpiops farkaci
        • Scorpiops tibetanus
      • Euscorpiops → (27 species)
        • Euscorpiops karschi
        • Euscorpiops lindnergi
        • Euscorpiops validus
        • Euscorpiops xui
        • Euscorpiops montanus
        • Euscorpiops shidian
        • Euscorpiops thaomischorum
        • Euscorpiops dakrong
        • Euscorpiops orioni
        • Euscorpiops yangi
        • Euscorpiops binghamii
        • Euscorpiops bhutanensis
        • Euscorpiops cavernicola
        • Euscorpiops beccaloniae
        • Euscorpiops kamengensis
        • Euscorpiops kaftani
        • Euscorpiops longimanus
        • Euscorpiops alexandreanneorum
        • Euscorpiops neradi
        • Euscorpiops puerensis
        • Euscorpiops asthenurus
        • Euscorpiops kubani
        • Euscorpiops sejnai
        • Euscorpiops novaki
        • Euscorpiops problematicus
        • Euscorpiops artemisae
        • Euscorpiops vachoni
    • Alloscorpiops → (6 species)
      • Alloscorpiops troglodytes
      • Alloscorpiops citadelle
      • Alloscorpiops wongpromi
      • Alloscorpiops calmonti
      • Alloscorpiops anthracinus
      • Alloscorpiops lindstroemii
    • Megacorminae → (8 species)
      • Megacormus → (5 species)
        • Megacormus segmentatus
        • Megacormus gertschi
        • Megacormus granosus
        • Megacormus grubbsi
        • Megacormus xichu
      • Plesiochactas → (3 species)
        • Plesiochactas mitchelli
        • Plesiochactas dilutus
        • Plesiochactas vasquezi
    • Troglocormus → (2 species)
      • Troglocormus willis
      • Troglocormus ciego
    • Neoscorpiops → (4 species)
      • Neoscorpiops maharashtraensis
      • Neoscorpiops deccanensis
      • Neoscorpiops satarensis
      • Neoscorpiops tenuicauda
    • Euscorpiinae → (50 species)
      • carpathicus species complex → (2 species)
        • Euscorpius carpathicus
        • Euscorpius concinnus
      • mingrelicus species complex → (4 species)
        • Euscorpius mingrelicus
        • Euscorpius gamma
        • Euscorpius phrygius
        • Euscorpius ciliciensis
      • Euscorpius sicanus
      • Euscorpius parthenopeius
      • Euscorpius balearicus
      • Euscorpius stahlavskyi
      • Euscorpius rahsenae
      • Euscorpius curcici
      • Euscorpius erymanthius
      • Euscorpius naupliensis
      • Euscorpius hadzii
      • Euscorpius beroni
      • Euscorpius flavicaudis
      • Euscorpius croaticus
      • Euscorpius avcii
      • Euscorpius koschewnikowi
      • Euscorpius aquilejensis
      • Euscorpius giachinoi
      • Euscorpius koci
      • Euscorpius alpha
      • Euscorpius yagmuri
      • Euscorpius uludagensis
      • Euscorpius deltshevi
      • Euscorpius drenskii
      • Euscorpius feti
      • Euscorpius arikani
      • Euscorpius mylonasi
      • Euscorpius italicus
      • Euscorpius tergestinus
      • Euscorpius koshewnikowi
      • Euscorpius kritscheri
      • Euscorpius celanus
      • Euscorpius oglasae
      • Crimean scorpion (Euscorpius tauricus)
      • Euscorpius solegladi
      • Euscorpius gocmeni
      • Euscorpius kinzelbachi
      • Euscorpius birulai
      • Euscorpius corcyraeus
      • Euscorpius honazicus
      • Euscorpius alanyaensis
      • Euscorpius vailatii
      • Euscorpius germanus
      • Euscorpius lycius
      • Euscorpius amorgensis
      • Euscorpius vignai
    • Parascorpiops montanus
    • Dasyscorpiops grandjeani
  • Superstitionia donensis

Other links

• Wikidata
• Wikipedia (language: ar)
• Encyclopedia of Life
• NCBI (genetics)"
• Open Tree of Life

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