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Base of the evolutionary tree for Dytiscoidea

Dytiscoidea (4578 species)
- unnamed group 1591162 → (4448 species)
  - predaceous diving beetles (Dytiscidae) → (4434 species)
    - unnamed group 1591918 → (3692 species)
      - unnamed group 1592750 → (2860 species)
      - unnamed group 1591919 → (832 species)
    - Hydronebrius → (4 species)
      - Hydronebrius cordaticollis
      - Hydronebrius kashmirensis
      - Hydronebrius mattheyi
      - Hydronebrius amplicollis
    - Heterhydrus → (5 species)
      - Heterhydrus sudanensis
      - Heterhydrus adipatus
      - Heterhydrus senegalensis
      - Heterhydrus ghanensis
      - Heterhydrus agaboides
    - Laccoporus → (2 species)
      - Laccoporus viator
      - Laccoporus nigritulus
    - Laccophilinae → (425 species)
      - Laccophilini → (423 species)
      - Agabetini → (2 species)
    - Hyphovatus → (3 species)
      - Hyphovatus prapatensis
      - Hyphovatus manfredi
      - Hyphovatus dismorphus
    - Liadytiscus → (3 species)
      - Liadytiscus cretaceus
      - Liadytiscus longitibialis
      - Liadytiscus latus
    - Colymbetinae → (143 species)
      - Colymbetini → (140 species)
      - Anisomeriini → (2 species)
      - Carabdytes upin
    - Laccomimus → (13 species)
      - Laccomimus bolivari
      - Laccomimus alvarengi
      - Laccomimus youngi
      - Laccomimus xikrin
      - Laccomimus pumilio
      - Laccomimus distinctus
      - Laccomimus bordoni
      - Laccomimus variegatus
      - Laccomimus amazonas
      - Laccomimus spinosus
      - Laccomimus malkini
      - Laccomimus improvidus
      - Laccomimus spangleri
    - Yolina → (10 species)
      - Yolina elegantula
      - Yolina royi
      - Yolina insignis
      - Yolina libera
      - Yolina kongouensis
      - Yolina chopardi
      - Yolina wewalkai
      - Yolina balfourbrownei
      - Yolina sima
      - Yolina baerti
    - Hemibidessus → (5 species)
      - Hemibidessus celinoides
      - Hemibidessus plaumanni
      - Hemibidessus spangleri
      - Hemibidessus spiroductus
      - Hemibidessus conicus
    - Japanolaccophilus → (2 species)
      - Japanolaccophilus niponensis
      - Japanolaccophilus beatificus
    - Tepuidessus → (2 species)
      - Tepuidessus breweri
      - Tepuidessus grulai
    - Neoclypeodytes → (24 species)
      - Neoclypeodytes nanus
      - Neoclypeodytes tumulus
      - Neoclypeodytes leachi
      - Neoclypeodytes amybethae
      - Neoclypeodytes megalus
      - Neoclypeodytes haroldi
      - Neoclypeodytes similis
      - Neoclypeodytes pictodes
      - Neoclypeodytes challeti
      - Neoclypeodytes edithae
      - Neoclypeodytes discretus
      - Neoclypeodytes roughleyi
      - Neoclypeodytes anasinus
      - Neoclypeodytes plicipennis
      - Neoclypeodytes ornatellus
      - Neoclypeodytes astrapus
      - Neoclypeodytes curtulus
      - Neoclypeodytes quadrinotatus
      - Neoclypeodytes cinctellus
      - Neoclypeodytes discedens
      - Neoclypeodytes latifrons
      - Neoclypeodytes fryii
      - Neoclypeodytes quadripustulatus
      - Neoclypeodytes obesus
    - Agnoshydrus → (8 species)
      - Agnoshydrus densus
      - Agnoshydrus schillhammeri
      - Agnoshydrus barong
      - Agnoshydrus ciampori
      - Agnoshydrus taiwanus
      - Agnoshydrus paulbrowni
      - Agnoshydrus laccophiloides
      - Agnoshydrus confusus
    - Africodytes → (5 species)
      - Africodytes rubromaculatus
      - Africodytes kongouensis
      - Africodytes maximus
      - Africodytes silvestris
      - Africodytes mopiensis
    - Lioporeus → (2 species)
      - Lioporeus pilatei
      - A Predaceous Diving Beetle (Lioporeus triangularis)
    - Andonectes → (13 species)
      - Andonectes similaris
      - Andonectes mildredae
      - Andonectes septentrionalis
      - Andonectes pineiroi
      - Andonectes intermedius
      - Andonectes milla
      - Andonectes apure
      - Andonectes meridensis
      - Andonectes trujillo
      - Andonectes maximus
      - Andonectes bordoni
      - Andonectes gregarius
      - Andonectes venezuelanus
    - Hydrodytinae → (3 species)
      - Hydrodytes inaciculatus
      - Hydrodytes opalinus
      - Hydrodytes dodgei
    - Mesoderus → (4 species)
      - Mesoderus ovatus
      - Mesoderus ventralis
      - Mesoderus punctatus
      - Mesoderus magnus
    - Lancetinae → (22 species)
      - Lancetes waterhousei
      - Lancetes delkeskampi
      - Lancetes borellii
      - Lancetes arauco
      - Lancetes tarsalis
      - Lancetes flavipes
      - Lancetes varius
      - Lancetes marginatus
      - Lancetes backstromi
      - Lancetes biremis
      - Lancetes subseriatus
      - Lancetes towianicus
      - Lancetes immarginatus
      - Lancetes flavoscutatus
      - Lancetes mixtus
      - Lancetes lanceolatus
      - Lancetes nigriceps
      - Lancetes praemorsus
      - Lancetes angusticollis
      - Lancetes falklandicus
      - Lancetes theresae
      - Lancetes dacunhae
    - Anginopachria → (3 species)
      - Anginopachria schoedli
      - Anginopachria ullrichi
      - Anginopachria prudeki
    - Platydytes → (4 species)
      - Platydytes incognitus
      - Platydytes coarctaticollis
      - Platydytes cooperae
      - Platydytes inspectatus
    - Geodessus → (2 species)
      - Geodessus besucheti
      - Geodessus kejvali
    - Phreatodessus → (2 species)
      - Phreatodessus hades
      - Phreatodessus pluto
    - Coptotominae → (7 species)
      - A Predaceous Diving Beetle (Coptotomus loticus)
      - Coptotomus lenticus
      - Coptotomus interrogatus
      - Coptotomus difficilis
      - A Predaceous Diving Beetle (Coptotomus longulus)
      - Coptotomus venustus
      - Coptotomus serripalpus
    - Morimotoa → (3 species)
      - Morimotoa morimotoi
      - Morimotoa phreatica
      - Morimotoa gigantea
    - Borneodessus zetteli
    - Liadyxianus kirejtshuki
    - Siettitia avenionensis
    - Dimitshydrus typhlops
    - Huxelhydrus syntheticus
    - Petrodessus conatus
    - Typhlodessus monteithi
    - Microhydrodytes elachistus
    - Sinoporus lineatus
    - Napodytes boki
    - Spanglerodessus shorti
    - Sinodytes hubbardi
    - Crinodessus amyae
    - Comal Springs Diving Beetle (Comaldessus stygius)
    - Trogloguignotus concii
    - Incomptodessus camachoi
    - Palaeogyrinus strigatus
    - Regimbartina pruinosa
    - Glessaria rostrata
    - Liadroporus elegans
    - Siamoporus deharvengi
    - Mesodytes rhantoides
    - Microdessus atomarius
  - unnamed group 1591163 → (14 species)
    - unnamed group 1591170 → (7 species)
      - Trout-stream beetle (Amphizoidae) → (5 species)
      - cliff water beetles (Aspidytidae) → (2 species)
    - diving beetles (Hygrobiidae) → (7 species)
- unnamed group 1591033 → (130 species)
  - Burrowing water beetle (Noteridae) → (129 species)
  - Meru phyllisae

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