Base of the evolutionary tree for Harpalus

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• Harpalus (475 species)
  • Omostropus minor
  • Omostropus kilimanus
  • Omostropus tersulus
  • Omostropus vicarius
  • Omostropus similis
  • Omostropus rotundatus
  • Omostropus cratognathoides
  • Omostropus bulirschi
  • Omostropus australis
  • Harpalus oustaleti
  • Harpalus distinguendus
  • Harpalus inconcinnus
  • Harpalus egorovi
  • Harpalus kiritshenkoi
  • Harpalus providens
  • Harpalus inexpectatus
  • Harpalus dissitus
  • A Ground Beetle (Harpalus indianus)
  • Harpalus lama
  • Harpalus apache
  • Harpalus parasinuatus
  • Harpalus fuscipennis
  • Harpalus janinae
  • Harpalus punctatostriatus
  • Harpalus pygmaeus
  • Harpalus triseriatus
  • Harpalus capicola
  • Harpalus hybridus
  • Harpalus rivalsi
  • Harpalus vittatus
  • Harpalus meghalayensis
  • Harpalus pulvinatus
  • Harpalus durangoensis
  • Harpalus famelicus
  • Harpalus disimuciulus
  • Harpalus mitridati
  • Harpalus imerinae
  • Harpalus massarti
  • Harpalus stoetznerianus
  • Harpalus kryzhanovskii
  • Harpalus abolitus
  • Harpalus flavescens
  • Harpalus litigiosus
  • Harpalus pulchrinulus
  • Harpalus vagans
  • Harpalus ingenuus
  • Harpalus rufomarginatus
  • Harpalus lampronotus
  • Harpalus minutulus
  • Harpalus bruckmanni
  • A Ground Beetle (Harpalus plenalis)
  • Harpalus rotundus
  • Harpalus angustipennis
  • A Carabid Beetle (Harpalus caliginosus)
  • Harpalus poncei
  • Harpalus venator
  • Harpalus roninus
  • Harpalus deletus
  • Harpalus neglectus
  • Harpalus oblitus
  • Harpalus defector
  • Harpalus fulvus
  • Harpalus wagneri
  • Harpalus caeruleatus
  • A Ground Beetle (Harpalus faunus)
  • Harpalus frater
  • Harpalus tenuestriatus
  • Harpalus knorri
  • Harpalus attenuatus
  • A Ground Beetle (Harpalus atrichatus)
  • Harpalus latus
  • Harpalus morvani
  • Harpalus prosperus
  • Harpalus simplicidens
  • Harpalus tinctulus
  • Harpalus sinuatipennis
  • Harpalus forsstroemi
  • Harpalus tithonus
  • Harpalus brachypterus
  • A Ground Beetle (Harpalus fulvilabris)
  • Harpalus subaeneus
  • Harpalus pallipes
  • Harpalus sanctaehelenae
  • Harpalus katiae
  • Harpalus kailiensis
  • Harpalus major
  • Harpalus seyrigi
  • Harpalus quadratus
  • Harpalus balli
  • Harpalus ganssuensis
  • Harpalus fulvipennis
  • Harpalus kabakianus
  • Harpalus optabilis
  • Harpalus aterrimus
  • Harpalus quenselii
  • Harpalus kmecoi
  • Harpalus subcylindricus
  • A Ground Beetle (Harpalus herbivagus)
  • A Ground Beetle (Harpalus ventralis)
  • Harpalus glasunovi
  • Harpalus petri
  • Harpalus dispar
  • A Ground Beetle (Harpalus fulgens)
  • Harpalus procognatus
  • Harpalus coreanus
  • Harpalus amplicollis
  • Harpalus melaneus
  • Harpalus singularis
  • Harpalus progrediens
  • Harpalus raphaili
  • Harpalus burmeisteri
  • Harpalus exiguus
  • Harpalus vernalis
  • Harpalus smyrnensis
  • Harpalus convexus
  • Harpalus tabidus
  • Harpalus madagascariensis
  • A Ground Beetle (Harpalus ochropus)
  • Harpalus obtusiusculus
  • Harpalus puetzi
  • Harpalus stygius
  • Harpalus anxius
  • Harpalus disaogashimensis
  • Harpalus stierlini
  • Harpalus sushenicus
  • Harpalus lateralis
  • Harpalus hirtipes
  • Harpalus servus
  • Harpalus parvulus
  • Harpalus aureus
  • Harpalus natalensis
  • Harpalus immixtus
  • Harpalus zhdankoi
  • Harpalus nero
  • Harpalus breviusculus
  • Harpalus spadiceus
  • Harpalus cautus
  • Harpalus xinjiangensis
  • Harpalus impressicollis
  • Harpalus jeanneli
  • Harpalus alexeevi
  • Harpalus masoreoides
  • Harpalus adenticulatus
  • Harpalus brunnipes
  • Harpalus nuperus
  • Harpalus tiridates
  • Harpalus davidi
  • Harpalus saxicola
  • Harpalus corrugatus
  • Harpalus marginellus
  • Harpalus kadyrbekovi
  • Harpalus ussuricus
  • Harpalus gilgil
  • Harpalus chobautianus
  • Harpalus stephani
  • A Ground Beetle (Harpalus indigens)
  • Harpalus farkaci
  • Harpalus babai
  • Harpalus tardus
  • Harpalus pseudohauserianus
  • Harpalus italus
  • Harpalus euchlorus
  • Harpalus nanniscus
  • Harpalus aogashimensis
  • Harpalus albanicus
  • A Ground Beetle (Harpalus rufibarbis)
  • Harpalus rubefactus
  • Harpalus cyclogonus
  • Harpalus araraticus
  • Harpalus vicarius
  • A Ground Beetle (Harpalus reversus)
  • Harpalus vernicosus
  • A Ground Beetle (Harpalus rubripes)
  • Harpalus fulvicornis
  • Harpalus chasanensis
  • Harpalus suturangulus
  • Harpalus foveiger
  • Harpalus basilewskyi
  • Harpalus honestus
  • Harpalus longipalmatus
  • Harpalus punctipennis
  • Harpalus nevadensis
  • Harpalus caiphus
  • Harpalus puncticeps
  • Harpalus sericatus
  • Harpalus subtruncatus
  • Harpalus turmalinus
  • Harpalus fuscoaeneus
  • Harpalus demelti
  • Harpalus udege
  • Harpalus oodioides
  • Harpalus kunarensis
  • Harpalus praticola
  • Harpalus periglabellus
  • Harpalus mariae
  • A Ground Beetle (Harpalus megacephalus)
  • Strawberry Seed Beetle (Harpalus rufipes)
  • Harpalus cilihumerus
  • Harpalus scabripectus
  • A Ground Beetle (Harpalus compar)
  • Harpalus dubius
  • Harpalus dilatatus
  • Harpalus lumbaris
  • Harpalus alpivagus
  • Harpalus aequicollis
  • A Ground Beetle (Harpalus opacipennis)
  • Harpalus politus
  • Harpalus ferghanensis
  • Harpalus platynotus
  • Harpalus spurius
  • Harpalus poussereaui
  • Harpalus chalcopterus
  • Harpalus kaznakovi
  • Harpalus aeneipennis
  • Harpalus manas
  • Harpalus spretus
  • Harpalus indicola
  • Harpalus kazanensis
  • Harpalus giacomazzoi
  • Harpalus potanini
  • Harpalus miles
  • Harpalus anatolicus
  • Harpalus retractus
  • Harpalus desertus
  • Brush-thighed Seed-eater (Harpalus froelichii)
  • Harpalus sulphuripes
  • Harpalus koshiensis
  • Harpalus arcuatus
  • Harpalus jordanus
  • Harpalus picipennis
  • Harpalus danieli
  • Harpalus cohni
  • Harpalus kandaharensis
  • Harpalus protractus
  • Harpalus circumpunctatus
  • Harpalus ulamaeformis
  • Harpalus metarsius
  • A Ground Beetle (Harpalus obnixus)
  • Harpalus salinulus
  • Harpalus amarellus
  • Harpalus decipiens
  • Harpalus maceratus
  • Harpalus franzi
  • Harpalus cardoni
  • Harpalus bucharicus
  • A Ground Beetle (Harpalus cordifer)
  • Harpalus hypogeomysis
  • Harpalus somnulentus
  • Harpalus pallidipennis
  • Harpalus ascetes
  • Harpalus fimetarius
  • Harpalus indicus
  • Harpalus pigmaeus
  • Harpalus diversicollis
  • Harpalus meridianus
  • Harpalus pseudoserripes
  • Harpalus xanthopus
  • Harpalus azumai
  • Harpalus lethierryi
  • Harpalus reitteri
  • Harpalus wohlberedti
  • Harpalus cyanopterus
  • Harpalus karamani
  • Harpalus suensoni
  • Harpalus turcicus
  • Harpalus tsushimanus
  • Harpalus idiotus
  • Harpalus cordatus
  • Harpalus kagyzmanicus
  • Harpalus longihornus
  • Harpalus kibonoti
  • Harpalus ussuriensis
  • Harpalus pseudophonoides
  • Harpalus lopezi
  • Harpalus ellipsis
  • Harpalus caspius
  • Harpalus strenuus
  • Harpalus rufitarsoides
  • Harpalus uniformis
  • Harpalus klapperichi
  • Harpalus conditus
  • Harpalus remboides
  • Harpalus redivivus
  • Harpalus sinis
  • Harpalus innocuus
  • Harpalus akinini
  • Harpalus torridoides
  • Harpalus tjanschanicus
  • Harpalus aenigma
  • Harpalus kozlovi
  • Harpalus niigatanus
  • Harpalus obliquus
  • Harpalus atripes
  • Harpalus nigrans
  • Harpalus anisodactyliformis
  • Harpalus nitens
  • Harpalus gregoryi
  • Harpalus salinus
  • Harpalus agnatus
  • Harpalus pleistocenicus
  • Harpalus colchicus
  • Harpalus angustitarsis
  • Harpalus szalliesi
  • Harpalus vanemdeni
  • Harpalus pecinai
  • Harpalus polyglyptus
  • Harpalus viridicupreus
  • Harpalus calathoides
  • Harpalus luteicornis
  • Harpalus angulatus
  • Harpalus excavatus
  • Harpalus gisellae
  • Harpalus chalcentus
  • Harpalus pseudotinctulus
  • Harpalus calceatus
  • Harpalus tichonis
  • Harpalus flavicornis
  • Harpalus ewaldi
  • Harpalus tardigradus
  • Harpalus grilli
  • Harpalus hatchi
  • Harpalus erosus
  • Harpalus chengjiangensis
  • Harpalus egenus
  • Harpalus corporosus
  • Harpalus rougemonti
  • A Ground Beetle (Harpalus pensylvanicus)
  • A Ground Beetle (Harpalus animosus)
  • Harpalus celax
  • Harpalus whitfieldi
  • Harpalus actiosus
  • Harpalus brevis
  • Harpalus reflexus
  • Harpalus pachys
  • Harpalus bitinctus
  • Harpalus stevenii
  • Harpalus somereni
  • Harpalus setiporus
  • Harpalus huachuca
  • Harpalus impressus
  • Harpalus serripes
  • Harpalus modestus
  • A Ground Beetle (Harpalus solitaris)
  • Harpalus makhekensis
  • Harpalus heeri
  • Harpalus cyrenaicus
  • Harpalus michailovi
  • Harpalus liasinus
  • Harpalus lewisii
  • Harpalus basuto
  • Harpalus inexspectatus
  • Harpalus hospes
  • Harpalus kadleci
  • Harpalus fukiensis
  • Harpalus nyassicus
  • Harpalus tangutorum
  • Harpalus australasiae
  • Harpalus pilosus
  • Harpalus hauserianus
  • Harpalus anxioides
  • Harpalus antonowi
  • Harpalus microdemas
  • Harpalus vereschaginae
  • Harpalus meteorus
  • Harpalus lederi
  • Harpalus asemus
  • Harpalus mauritanicus
  • Harpalus crates
  • Harpalus basanicus
  • Harpalus diligens
  • Harpalus kirgisicus
  • Harpalus bungii
  • A Ground Beetle (Harpalus erythropus)
  • Harpalus lugubris
  • Harpalus bellieri
  • Harpalus eous
  • Harpalus macronotus
  • Harpalus ovtshinnikovi
  • Harpalus michaili
  • Harpalus aesculanus
  • Harpalus semenowi
  • Harpalus compressus
  • Harpalus trichophorus
  • Harpalus texanus
  • Harpalus sinuatus
  • Harpalus natalicus
  • Harpalus pusillus
  • Harpalus geniculatus
  • Harpalus tridens
  • Harpalus mairei
  • Harpalus paratus
  • Harpalus agilis
  • A Ground Beetle (Harpalus laticeps)
  • Harpalus alexandrae
  • Harpalus cupreus
  • Harpalus kabakovi
  • Harpalus microthorax
  • A Ground Beetle (Harpalus fraternus)
  • Harpalus chrysopus
  • A Ground Beetle (Harpalus fuscipalpis)
  • Harpalus metallinus
  • Harpalus smaragdinus
  • Harpalus muciulus
  • Harpalus ebeninus
  • Harpalus plancyi
  • A Ground Beetle (Harpalus nigritarsis)
  • Harpalus pastor
  • Harpalus cephalotes
  • Harpalus griseus
  • Harpalus autumnalis
  • Harpalus terrestris
  • Harpalus chiloschizontus
  • Harpalus amariformis
  • Harpalus discrepans
  • Harpalus contemptus
  • Harpalus baleensis
  • Harpalus karakorum
  • Harpalus constrictus
  • Harpalus tibeticus
  • Harpalus veterum
  • Harpalus acupalpoides
  • Harpalus schlotheimi
  • Harpalus ulomaeformis
  • Harpalus siculus
  • Harpalus heyrovskyi
  • Harpalus rumelicus
  • Harpalus gravis
  • Harpalus anactus
  • Harpalus martini
  • Harpalus alajensis
  • Harpalus pumilus
  • Harpalus wadiensis
  • Harpalus medvedevi
  • Harpalus aeneus
  • Harpalus arnoldii
  • A Ground Beetle (Harpalus affinis)
  • Harpalus atratus
  • Harpalus tarsalis
  • Harpalus sinicus
  • Harpalus mlynari
  • Harpalus yinchuanensis
  • A Ground Beetle (Harpalus laevipes)
  • Harpalus praecurrens
  • Harpalus jureceki
  • Harpalus cisteloides
  • Harpalus torosensis
  • Harpalus badakschanus
  • Harpalus penglainus
  • Harpalus sundaicus
  • Harpalus agakhaniantzi
  • Harpalus numidicus
  • Harpalus silipes
  • Harpalus zabroides
  • A Ground Beetle (Harpalus amputatus)
  • Harpalus hartmanni
  • Harpalus davidianus
  • Harpalus fuscicornis
  • Harpalus beneshi
  • Harpalus corpulentus
  • Harpalus capito
  • Harpalus rufipalpis
  • Harpalus fokienensis
  • Harpalus pterostichus
  • Harpalus lheritieri
  • Harpalus rufiscapus
  • Harpalus erraticus
  • Harpalus dimidiatus
  • Harpalus lutshniki
  • Harpalus laetus
  • Harpalus stricticollis
  • Harpalus obtusus
  • Harpalus viridanus
  • Harpalus alienus
  • Harpalus taciturnus
  • Harpalus liobasis

Other links

• Wikidata
• Wikipedia (language: en)
• Wikipedia (language: es)
• Wikipedia (language: fr)
• Wikipedia (language: ru)
• Wikipedia (language: pl)
• Wikipedia (language: nl)
• Wikipedia (language: tr)
• Wikipedia (language: sv)
• Wikipedia (language: vi)
• Encyclopedia of Life
• NCBI (genetics)"
• Open Tree of Life

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