Base of the evolutionary tree for Leucosioidea

← parent

• Leucosioidea (603 species)
  • purse crabs (Leucosiidae) → (596 species)
    • Ebaliinae → (472 species)
      • Arcania → (24 species)
        • Arcania erinacea
        • Arcania echinata
        • Arcania duodecimspinosa
        • Arcania cornuta
        • Arcania novemspinosa
        • Arcania marinduquensis
        • Arcania shuishuni
        • Arcania globata
        • Arcania elongata
        • Arcania tropicalis
        • Arcania fungilifera
        • Arcania undecimspinosa
        • Arcania tuberculata
        • Arcania heptacantha
        • Arcania muricata
        • Arcania granulipes
        • Arcania foliolata
        • Arcania gracilis
        • Arcania aspera
        • Arcania orientalis
        • Arcania septemspinosa
        • Arcania sagamiensis
        • Arcania cornigera
        • Arcania brevifrons
      • Lyphira → (5 species)
        • Lyphira ovata
        • Lyphira natalensis
        • Lyphira heterograna
        • Lyphira perplexa
        • Lyphira georgei
      • Pyrhila → (3 species)
        • Pyrhila pisum
        • Pyrhila biprotubera
        • Pyrhila carinata
      • Pseudophilyra → (18 species)
        • Pseudophilyra punctulata
        • Pseudophilyra granulimarginata
        • Pseudophilyra burmensis
        • Pseudophilyra pusilla
        • Pseudophilyra melita
        • Pseudophilyra albimaculata
        • Pseudophilyra perryi
        • Pseudophilyra blandfordi
        • Pseudophilyra burmaensis
        • Pseudophilyra elongatella
        • Pseudophilyra parilis
        • Pseudophilyra polita
        • Pseudophilyra tenuipes
        • Pseudophilyra intermedia
        • Pseudophilyra woodmasoni
        • Pseudophilyra tridentata
        • Pseudophilyra deficiens
        • Pseudophilyra nanshaensis
      • Uhlias → (2 species)
        • eroded clutch crab (Uhlias limbatus)
        • Uhlias ellipticus
      • Randallia → (15 species)
        • Randallia laevis
        • Randallia jingomao
        • Randallia americana
        • Randallia saitoensis
        • Randallia angelica
        • Randallia curacaoensis
        • Randallia granulata
        • Randallia strouhali
        • Randallia agaricias
        • globose sand crab (Randallia ornata)
        • Randallia minuta
        • Randallia gilberti
        • Randallia pleistocenica
        • Randallia prolanata
        • Randallia bulligera
      • Myra → (27 species)
        • Myra australis
        • Myra subcarinata
        • Myra hainanica
        • Myra eudactylus
        • Myra trispinosa
        • Myra tumidospina
        • Myra fugax
        • Myra brevimana
        • Myra digitata
        • Myra curtimana
        • Myra pernix
        • Myra subgranulata
        • Myra elegans
        • Myra wenjii
        • Myra brevisulcata
        • Myra emarginata
        • Myra celeris
        • Myra arachnoides
        • Myra affinis
        • Myra currax
        • Myra intermedia
        • Myra paenitensis
        • Myra mammilaris
        • Myra grandis
        • Myra amoena
        • Myra fijiensis
        • Myra biconica
      • Toru → (5 species)
        • Toru granuloides
        • Toru trituberculatus
        • Toru pilus
        • Toru septimus
        • Toru mesjatzevi
      • Ancylodactyla → (3 species)
        • Ancylodactyla elata
        • Ancylodactyla nana
        • Ancylodactyla elongata
      • Lithadia → (8 species)
        • Lithadia barnardi
        • Lithadia conica
        • Lithadia vertiginosa
        • carinate clutch crab (Lithadia cadaverosa)
        • Lithadia brasiliensis
        • Lithadia cumingii
        • Lithadia obliqua
        • Lithadia granulosa
      • Ryphila → (3 species)
        • Ryphila verrucosa
        • Ryphila cancellus
        • Ryphila bertrandi
      • Nucia → (13 species)
        • Nucia speciosa
        • Nucia fennemai
        • Nucia borneoensis
        • Nucia miliaris
        • Nucia rosea
        • Nucia ingens
        • Nucia calculoides
        • Nucia coxi
        • Nucia platyspinosa
        • Nucia tuberculosa
        • Nucia pulchella
        • Nucia baripadensis
        • Nucia bouvieri
      • Ihleus → (2 species)
        • Ihleus lanatus
        • Ihleus villosus
      • Leucosilia → (3 species)
        • Leucosilia bananensis
        • Leucosilia jurinii
        • Leucosilia maldivensis
      • Paranursia → (2 species)
        • Paranursia abbreviata
        • Paranursia acharyai
      • Galilia → (2 species)
        • Galilia narusei
        • Galilia petricola
      • Nursilia → (3 species)
        • Nursilia sinica
        • Nursilia tonsor
        • Nursilia dentata
      • Persephona → (18 species)
        • Persephona prepunctata
        • Persephona edwardsii
        • Persephona townsendi
        • Persephona rodesae
        • Persephona manningi
        • Persephona enigmatica
        • Persephona lichtenstenii
        • Persephona punctata
        • Persephona orbicularis
        • mottled purse crab (Persephona mediterranea)
        • Persephona finneganae
        • Persephona bigranulata
        • Persephona niemeyeri
        • Persephona subovata
        • Persephona aquilonaris
        • Persephona lichtensteinii
        • Persephona invalida
        • pink purse crab (Persephona crinita)
      • Tokoyo → (3 species)
        • Tokoyo trinuclodea
        • Tokoyo eburnea
        • Tokoyo cirrata
      • Raylilia → (5 species)
        • Raylilia coniculifera
        • Raylilia gracilipes
        • Raylilia intermedia
        • Raylilia uenoi
        • Raylilia mirabilis
      • Merocryptoides → (3 species)
        • Merocryptoides peteri
        • Merocryptoides ohtsukai
        • Merocryptoides frontalis
      • Nobiliella → (3 species)
        • Nobiliella margaritata
        • Nobiliella jousseaumei
        • Nobiliella cornigera
      • Nursia → (21 species)
        • Nursia guinotae
        • Nursia lamellata
        • Nursia dimorpha
        • Nursia plicata
        • Nursia tohae
        • Nursia alata
        • Nursia phylloides
        • Nursia rhomboidalis
        • Nursia weberi
        • Nursia trilobata
        • Nursia nasuta
        • Nursia sinuata
        • Nursia persica
        • Nursia elegans
        • Nursia japonica
        • Nursia minor
        • Nursia sexangulata
        • Nursia lar
        • Nursia blandfordi
        • Nursia mimetica
        • Nursia hamipleopoda
      • Heteronucia → (18 species)
        • Heteronucia laminata
        • Heteronucia tuberculata
        • Heteronucia spinifera
        • Heteronucia perlata
        • Heteronucia granulata
        • Heteronucia elegans
        • Heteronucia fujitai
        • Heteronucia xincunensis
        • Heteronucia mesanensis
        • Heteronucia vesiculosa
        • Heteronucia toyoshioae
        • Heteronucia minuta
        • Heteronucia venusta
        • Heteronucia obfastigiatus
        • Heteronucia oeschi
        • Heteronucia globata
        • Heteronucia margaritata
        • Heteronucia angulata
      • Oreotlos → (12 species)
        • Oreotlos etor
        • Oreotlos angulatus
        • Oreotlos latus
        • Oreotlos potanus
        • Oreotlos heuretos
        • Oreotlos bertrandi
        • Oreotlos encymus
        • Oreotlos speciosus
        • Oreotlos pala
        • Oreotlos havelocki
        • Oreotlos lagarodes
        • Oreotlos pax
      • Iliacantha → (7 species)
        • shouldered purse crab (Iliacantha sparsa)
        • Iliacantha globosa
        • longfinger purse crab (Iliacantha subglobosa)
        • Iliacantha panamanica
        • Iliacantha hancocki
        • Iliacantha schmitti
        • Iliacantha liodactylus
      • Merocryptus → (5 species)
        • Merocryptus altavillensis
        • Merocryptus lambriformis
        • Merocryptus boletisculpta
        • Merocryptus boletifer
        • Merocryptus obsoletus
      • Ixa → (9 species)
        • Ixa holthuisi
        • Ixa edwardsii
        • Ixa monodi
        • Ixa cylindrus
        • Ixa profundus
        • Ixa acuta
        • Ixa inermis
        • Ixa pulcherrima
        • Ixa investigatoris
      • Myrine → (2 species)
        • Myrine acutidens
        • Myrine kessleri
      • Philyra → (46 species)
        • Philyra granigera
        • Philyra olivacea
        • Philyra alveola
        • Philyra scabriuscula
        • Philyra alcocki
        • Philyra nishihirai
        • Philyra samia
        • Philyra rudis
        • Philyra plana
        • Philyra fuliginosa
        • Philyra syndactyla
        • Philyra trusanensis
        • Philyra taekoae
        • Philyra collinsi
        • Philyra pusilla
        • Philyra sexangula
        • Philyra hayasakai
        • Philyra cranium
        • Philyra malefactrix
        • Philyra unidentata
        • Philyra angularis
        • Philyra cancella
        • Philyra macrophthalma
        • Philyra rectangularis
        • Philyra miyamotoi
        • Philyra porcellanea
        • Philyra sagittifera
        • Philyra tridentata
        • Philyra marginata
        • Philyra hormozganensis
        • Philyra concinnus
        • Philyra adamsii
        • Philyra ferrica
        • Philyra kanekoi
        • Philyra corallicola
        • Philyra bicornis
        • Philyra shihcheni
        • Philyra tanakai
        • Philyra nishimotoi
        • Philyra iriomotensis
        • Philyra globus
        • Philyra orbicularis
        • Philyra misagoana
        • Philyra scabra
        • Philyra zhoushanensis
        • Philyra platychira
      • Ilia → (4 species)
        • nut-crab (Ilia nucleus)
        • Ilia spinosa
        • Ilia pliocenica
        • Ilia pliocaenica
      • Ebalia → (83 species)
        • Ebalia intermedia
        • Ebalia stellaris
        • Ebalia glans
        • sculptured clutch crab (Ebalia cariosa)
        • Ebalia quadridentata
        • Ebalia rotundata
        • Ebalia sakaii
        • Ebalia maldivensis
        • Ebalia vanstraeleni
        • Ebalia philippinensis
        • Ebalia hungarica
        • Ebalia fragifera
        • Ebalia nana
        • Ebalia tosaensis
        • Ebalia edwardsii
        • Ebalia nobilii
        • Ebalia dimorphoides
        • Ebalia meulenkampi
        • Ebalia conifera
        • Ebalia lambriformis
        • Ebalia orientalis
        • Ebalia serenei
        • Ebalia globulosa
        • Ebalia koyo
        • Ebalia scabriuscula
        • Ebalia glomus
        • Ebalia spinifera
        • Ebalia ferganensis
        • Ebalia agglomus
        • Ebalia diadumena
        • Ebalia tuberculosa
        • Ebalia takedai
        • Ebalia quadrata
        • Ebalia heterochalaza
        • Ebalia elegans
        • Ebalia clarionensis
        • Ebalia longispinosa
        • Ebalia webberi
        • Ebalia lacertosa
        • Ebalia longimana
        • Ebalia magdalenensis
        • Ebalia affinis
        • Ebalia laevis
        • Ebalia dentifrons
        • Ebalia brevimana
        • Ebalia discrepans
        • Ebalia granulosa
        • Ebalia braminae
        • Ebalia multiangulata
        • Ebalia punctulata
        • Ebalia lamarmorai
        • Ebalia humilis
        • Ebalia ypsilon
        • Ebalia woodmasoni
        • thinarm clutch crab (Ebalia stimpsoni)
        • Ebalia jacqueshermani
        • Pennant's nut crab (Ebalia tuberosa)
        • Ebalia nudipes
        • Ebalia vahldieki
        • Ebalia sculpta
        • Ebalia hayamaensis
        • Ebalia ramsayi
        • Ebalia pondoensis
        • Ebalia granulata
        • Bryer's nut crab (Ebalia tumefacta)
        • Ebalia yokoyai
        • Ebalia salamensis
        • Ebalia nux
        • Ebalia scandens
        • Ebalia hancocki
        • Ebalia crassipes
        • Ebalia bituberculata
        • Ebalia rhomboidalis
        • Ebalia oersi
        • Ebalia cryptocnemoides
        • Ebalia cristata
        • Ebalia abdominalis
        • Ebalia postulans
        • Ebalia barnardi
        • Ebalia kakegawaensis
        • Ebalia jordani
        • Ebalia fucinii
        • Cranch's nut crab (Ebalia cranchii)
      • Alox → (14 species)
        • Alox somphos
        • Alox rugosum
        • Alox antheos
        • Alox zalion
        • Alox naispela
        • Alox tormos
        • Alox ornatum
        • Alox latusoides
        • Alox glene
        • Alox liklik
        • Alox chaunos
        • Alox uru
        • Alox bothros
        • Alox patella
      • Praebebalia → (15 species)
        • Praebebalia kumanoensis
        • Praebebalia madagascariensis
        • Praebebalia mosakiana
        • Praebebalia nanhaiensis
        • Praebebalia longidactyla
        • Praebebalia extensiva
        • Praebebalia magna
        • Praebebalia fasciata
        • Praebebalia dondonae
        • Praebebalia fungifera
        • Praebebalia taeniata
        • Praebebalia fujianensis
        • Praebebalia pisiformis
        • Praebebalia septemspinosa
        • Praebebalia sikokuensis
      • Bellidilia → (3 species)
        • Bellidilia undecimspinosa
        • Bellidilia cheesmani
        • Bellidilia laevis
      • Urashima → (2 species)
        • Urashima pustuloides
        • Urashima lamellidentatus
      • Ebaliopsis → (2 species)
        • Ebaliopsis erosa
        • Ebaliopsis viadieri
      • Speloeophorus → (10 species)
        • Speloeophorus schmitti
        • Speloeophorus brasiliensis
        • smallhole clutch crab (Speloeophorus pontifer)
        • Speloeophorus microspeos
        • Speloeophorus callapoides
        • Speloeophorus inflatus
        • Speloeophorus subcircularis
        • Speloeophorus digueti
        • twohole clutch crab (Speloeophorus nodosus)
        • bighole clutch crab (Speloeophorus elevatus)
      • Tanaoa → (9 species)
        • Tanaoa kuka
        • Tanaoa nanus
        • Tanaoa granulatus
        • Tanaoa granulosa
        • Tanaoa distinctus
        • Tanaoa serenei
        • Tanaoa retpela
        • Tanaoa pustulosus
        • Tanaoa speciosus
      • Hiplyra → (9 species)
        • Hiplyra michellinae
        • Hiplyra varietgata
        • Hiplyra sagitta
        • Hiplyra platycheir
        • Hiplyra variegata
        • Hiplyra elegans
        • Hiplyra ramil
        • Hiplyra longimana
        • Hiplyra ramli
      • Parilia → (5 species)
        • Parilia tuberculata
        • Parilia major
        • Parilia ovata
        • Parilia pattersoni
        • Parilia alcocki
      • Oreophorus → (7 species)
        • Oreophorus fenestrus
        • Oreophorus speciosus
        • Oreophorus alcicornis
        • Oreophorus reticulatus
        • Oreophorus crosnieri
        • Oreophorus holthuisi
        • Oreophorus horridus
      • granulose purse crab (Acanthilia intermedia)
      • Pseudomyra mbizi
      • Atlantolocia laevidorsalis
      • Dolos petraeus
      • Favus granulatus
      • Nuciops modestus
      • Atlantophila cristata
      • Coralliocryptus caementa
      • spurfinger purse crab (Callidactylus asper)
      • Atlantotlos rhombifer
      • Afrophila punctata
      • Speloeophoroides capixaba
      • Kabutos durandi
      • Heterolithadia fallax
      • Orientotlos iishibai
      • Praosia punctata
      • Tlos muriger
      • fivespine purse crab (Myropsis quinquespinosa)
      • Cateios frontalis
    • Cryptocneminae → (27 species)
      • Cryptocnemus → (21 species)
        • Cryptocnemus obolus
        • Cryptocnemus holdsworthi
        • Cryptocnemus macrognathus
        • Cryptocnemus crenulatus
        • Cryptocnemus mortenseni
        • Cryptocnemus vincentianus
        • Cryptocnemus trapezoides
        • Cryptocnemus calmani
        • Cryptocnemus hemispheroides
        • Cryptocnemus siamensis
        • Cryptocnemus chinensis
        • Cryptocnemus aberrans
        • Cryptocnemus haddoni
        • Cryptocnemus pentagonus
        • Cryptocnemus marginatus
        • Cryptocnemus stimpsoni
        • Cryptocnemus tuberosus
        • Cryptocnemus grandidieri
        • Cryptocnemus kamekii
        • Cryptocnemus trigonus
        • Cryptocnemus planus
      • Leucisca → (4 species)
        • Leucisca squalina
        • Leucisca levigena
        • Leucisca rubifera
        • Leucisca halimedophila
      • Lissomorpha haswelli
      • Onychomorpha lamelligera
    • Leucosiinae → (97 species)
      • Leucosia → (38 species)
        • Leucosia calcarata
        • Leucosia margaritaria
        • Leucosia serenei
        • Leucosia tricarinata
        • Leucosia jecusculum
        • Leucosia ovalata
        • Leucosia whitmeei
        • Leucosia tetraodon
        • Leucosia compressa
        • Leucosia margaritacea
        • Leucosia haswelli
        • Leucosia rubripalma
        • Leucosia leslii
        • Leucosia tutongensis
        • Leucosia taiwanica
        • Leucosia phyllocheira
        • Leucosia anatum
        • Leucosia brevior
        • Leucosia reticulata
        • Leucosia brevimana
        • Leucosia longiangulata
        • Leucosia formosensis
        • Leucosia laevimana
        • Leucosia martini
        • Leucosia subrhomboidalis
        • Leucosia prevostiana
        • Leucosia longimaculata
        • Leucosia subrhomboidea
        • Leucosia sima
        • Leucosia ocellata
        • Leucosia craniolaris
        • Leucosia moresbiensis
        • Leucosia punctata
        • Leucosia longibrachia
        • Leucosia affinis
        • Leucosia corallicola
        • Leucosia persica
        • Leucosia takamii
      • Seulocia → (11 species)
        • Seulocia cristata
        • Seulocia rhomboidalis
        • Seulocia pubescens
        • Seulocia latirostrata
        • Seulocia crepuscula
        • Seulocia vittata
        • Seulocia laevimana
        • Seulocia anahita
        • Seulocia truncata
        • Seulocia gagulae
        • Seulocia pulchra
      • Urnalana → (24 species)
        • Urnalana parhaematostica
        • Urnalana purarensis
        • Urnalana chevreti
        • Urnalana haematosticta
        • Urnalana pulchella
        • Urnalana elatoides
        • Urnalana minuta
        • Urnalana thysanotus
        • Urnalana foresti
        • Urnalana cristata
        • Urnalana cumingii
        • Urnalana chevertii
        • Urnalana angulata
        • Urnalana granulimera
        • Urnalana elatula
        • Urnalana flammea
        • Urnalana hilaris
        • Urnalana margaritata
        • Urnalana insularis
        • Urnalana erythema
        • Urnalana whitei
        • Urnalana elata
        • Urnalana hirsuta
        • Urnalana parahaematostica
      • Euclosiana → (13 species)
        • Euclosia obtusifrons
        • Euclosia unidentata
        • Euclosiana guinotae
        • Euclosiana scitula
        • Euclosiana vella
        • Euclosiana concinna
        • Euclosiana unidentata
        • Euclosiana tornatilia
        • Euclosiana rotundifrons
        • Euclosiana exquisita
        • Euclosiana crosnieri
        • Euclosiana obtusifrons
        • Euclosiana nitida
      • Soceulia → (4 species)
        • Soceulia alainia
        • Soceulia marmorea
        • Soceulia brunnea
        • Soceulia major
      • Coleusia → (7 species)
        • Coleusia signata
        • Coleusia huilianae
        • Coleusia janani
        • Coleusia rangita
        • Coleusia urania
        • Coleusia biannulata
        • Coleusia magna
  • Iphiculidae → (7 species)
    • Pariphiculus → (4 species)
      • Pariphiculus agariciferus
      • Pariphiculus coronatus
      • Pariphiculus mariannae
      • Pariphiculus stellatus
    • Iphiculus → (3 species)
      • Iphiculus spongiosus
      • Iphiculus eliasi
      • Iphiculus convexus

Other links

• Wikidata
• Wikipedia (language: en)
• Wikipedia (language: fr)
• Wikipedia (language: zh)
• Wikipedia (language: nl)
• Wikipedia (language: fa)
• Wikipedia (language: ko)
• Encyclopedia of Life
• NCBI (genetics)"
• Open Tree of Life

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