Base of the evolutionary tree for Licininae

← parent

• Licininae (1352 species)
  • Chlaeniini → (1044 species)
    • Chlaenius → (1043 species)
      • Chlaenius simba
      • Chlaenius vestitus
      • Chlaenius eneides
      • Chlaenius natalensis
      • Chlaenius denticulatus
      • Chlaenius sangaicus
      • Chlaenius amabilis
      • Chlaenius scutellaris
      • Chlaenius madrasensis
      • Chlaenius obsidianoides
      • Chlaenius occultus
      • Chlaenius pugni
      • Chlaenius sellatus
      • Chlaenius pseudocupreolus
      • Chlaenius freyi
      • Chlaenius dondoensis
      • Chlaenius lobozianus
      • Chlaenius nyassaensis
      • Chlaenius africanicus
      • Chlaenius barkeri
      • Chlaenius nitidiceps
      • Chlaenius baehri
      • Chlaenius dianus
      • Chlaenius clarksoni
      • Chlaenius alternans
      • Chlaenius fuscicornis
      • Chlaenius nigricans
      • Chlaenius flavofemoratus
      • Chlaenius nyika
      • Chlaenius hornburgi
      • Chlaenius darlingensis
      • Chlaenius sykesi
      • Chlaenius bifenestratus
      • Chlaenius schatzmayri
      • Chlaenius indicus
      • Chlaenius obscuripennis
      • Chlaenius lativittis
      • Chlaenius tetracelis
      • Chlaenius sumbawanus
      • Chlaenius teani
      • Chlaenius jos
      • Chlaenius allacteus
      • Chlaenius stenoristus
      • Chlaenius ecuadoricus
      • Chlaenius claviger
      • Chlaenius deserticola
      • Chlaenius semiviridis
      • Chlaenius flaviguttatus
      • Chlaenius fallax
      • Chlaenius birmanicus
      • Chlaenius quadrisulcatus
      • Chlaenius virescens
      • Chlaenius insularis
      • Chlaenius episcopalis
      • Ground beetle (Chlaenius lithophilus)
      • Chlaenius subcostatus
      • Chlaenius decellei
      • Chlaenius pimalicus
      • Chlaenius holomelas
      • Chlaenius marginipennis
      • Chlaenius vietnamensis
      • Chlaenius haeckeli
      • Chlaenius lineicinctus
      • Chlaenius pauxillus
      • Chlaenius obscurus
      • Chlaenius juvencus
      • Chlaenius pubifer
      • Chlaenius jactus
      • Chlaenius pseudoraffrayi
      • Chlaenius parepus
      • Chlaenius luisae
      • Chlaenius flavipes
      • Chlaenius arakanensis
      • Chlaenius topali
      • Chlaenius vientianensis
      • Chlaenius carbonatus
      • Chlaenius balthasari
      • Chlaenius luluanus
      • Chlaenius similis
      • Chlaenius melancholicus
      • Chlaenius gansuensis
      • Chlaenius changwatensis
      • Chlaenius hutiaoxiaensis
      • Chlaenius limbipennis
      • Chlaenius oberthueri
      • Chlaenius rufomarginatus
      • Chlaenius grootfonteinensis
      • Chlaenius pailinensis
      • Chlaenius rebellis
      • Chlaenius tenuis
      • Chlaenius tansaniensis
      • Chlaenius leopoldi
      • Chlaenius amoenus
      • Chlaenius oxygonus
      • Chlaenius neochloodes
      • Chlaenius tetraphacus
      • Chlaenius maculiger
      • Chlaenius huedepohli
      • Chlaenius imperialis
      • Chlaenius longicornis
      • Chlaenius flavicornis
      • Chlaenius epicosmoides
      • Chlaenius circumductus
      • Chlaenius porinus
      • Chlaenius vinhphuensis
      • Chlaenius dalibaiensis
      • Chlaenius peringueyi
      • Chlaenius paenulatus
      • Chlaenius ammon
      • Chlaenius sparsepunctatus
      • Chlaenius gardoensis
      • Chlaenius acroxanthus
      • Chlaenius sierraleonensis
      • Chlaenius dhawalagiriensis
      • Chlaenius phaenoderus
      • Chlaenius cupripennis
      • Chlaenius nigrita
      • Chlaenius itombwanus
      • Chlaenius ophonoides
      • Chlaenius aurolimbatus
      • Chlaenius superbus
      • Chlaenius discicollis
      • Chlaenius heyrovskyi
      • Chlaenius litongaensis
      • Chlaenius pantarensis
      • Chlaenius buquetii
      • Chlaenius tanahrataensis
      • Chlaenius grundmanni
      • Chlaenius lomsakensis
      • Chlaenius darfurensis
      • Chlaenius leleupi
      • Chlaenius unyorensis
      • Chlaenius immunitus
      • Chlaenius suppletor
      • Chlaenius cupreopurpureus
      • Chlaenius mpanga
      • Chlaenius cumatilis
      • Chlaenius texanus
      • Chlaenius xanthospilus
      • Chlaenius xanthomerus
      • Chlaenius androyanus
      • Chlaenius circumdatus
      • Chlaenius femoratus
      • Chlaenius abjectus
      • Chlaenius sterbai
      • Chlaenius unicolor
      • Chlaenius paromius
      • Chlaenius bicolor
      • Ground beetle (Chlaenius sericeus)
      • Chlaenius lomii
      • Chlaenius rubricrus
      • Chlaenius roeschkei
      • Chlaenius notabilis
      • Chlaenius caeruleicollis
      • Chlaenius kabompo
      • Chlaenius langsonensis
      • Chlaenius schoenherri
      • Chlaenius tripustulatus
      • Chlaenius putzeysi
      • Chlaenius tingaudi
      • Chlaenius violaceipennis
      • Chlaenius amplians
      • Chlaenius virgulifer
      • Chlaenius feanus
      • Chlaenius adonis
      • Chlaenius alfredpuchneri
      • Chlaenius tudicus
      • Chlaenius sassanus
      • Chlaenius coscinioderus
      • Chlaenius aeratus
      • Chlaenius posticemaculatus
      • Chlaenius mederici
      • Chlaenius croyi
      • Chlaenius wittmerianus
      • Chlaenius bioculatus
      • Chlaenius tartagalensis
      • Chlaenius oneili
      • Chlaenius rufithorax
      • Chlaenius somereni
      • Chlaenius ruthmuellerae
      • Chlaenius variipes
      • Chlaenius surdipennis
      • Chlaenius anthraceus
      • Chlaenius geayi
      • Chlaenius morettoi
      • Chlaenius laosensis
      • Chlaenius porphyrius
      • Chlaenius duvaucelii
      • Chlaenius kindermanni
      • Chlaenius humeralis
      • Chlaenius chalcothorax
      • Chlaenius ampanihyensis
      • Chlaenius stschukini
      • Chlaenius rhysonotus
      • Chlaenius kapangae
      • Chlaenius agiloides
      • Chlaenius abstersus
      • Chlaenius apollo
      • Chlaenius cuprithorax
      • Chlaenius cubanus
      • Chlaenius purpureus
      • Chlaenius spathulifer
      • Chlaenius gotschii
      • Chlaenius coeruleipennis
      • Chlaenius breuningi
      • Chlaenius riparius
      • Chlaenius baxi
      • Chlaenius kashituensis
      • Chlaenius lederi
      • Chlaenius uluguruanus
      • Chlaenius obliquatus
      • Chlaenius insignis
      • Chlaenius pictus
      • Chlaenius similatus
      • Chlaenius tenellus
      • Chlaenius nossibianus
      • Chlaenius pecirkai
      • Chlaenius janus
      • Chlaenius moraveci
      • Ground beetle (Chlaenius pennsylvanicus)
      • Chlaenius validus
      • Chlaenius circumscriptus
      • Chlaenius keniaensis
      • Chlaenius submarginatus
      • Chlaenius pradieri
      • Chlaenius cyanipennis
      • Chlaenius jamaicae
      • Chlaenius maculiceps
      • Chlaenius loango
      • Chlaenius pretiosus
      • Chlaenius velocipes
      • Chlaenius borgouensis
      • Chlaenius koltzei
      • Chlaenius soccatus
      • Chlaenius dinodoides
      • Chlaenius ovalipennis
      • Chlaenius cinctus
      • Chlaenius bonelli
      • Chlaenius gonioderus
      • Chlaenius nepos
      • Chlaenius luteicauda
      • Chlaenius macropus
      • Chlaenius cruciatus
      • Chlaenius subferrugineus
      • Chlaenius vafer
      • Chlaenius laetiusculus
      • Chlaenius chagga
      • Chlaenius longulus
      • Chlaenius sulcipennis
      • Chlaenius frater
      • Chlaenius sobrinus
      • Chlaenius leucoscelis
      • Chlaenius scabricollis
      • Ground beetle (Chlaenius brevilabris)
      • Chlaenius sichuanensis
      • Chlaenius tristis
      • Chlaenius stygius
      • Chlaenius blanditus
      • Chlaenius rotundus
      • Chlaenius moheliensis
      • Chlaenius thieleni
      • Chlaenius nigricornis
      • Chlaenius costipennis
      • Chlaenius rajasthanensis
      • Chlaenius pulchriceps
      • Chlaenius kraatzi
      • Chlaenius alluaudianus
      • Chlaenius azurescens
      • Chlaenius puberulus
      • Chlaenius lafertei
      • Chlaenius diephelus
      • Chlaenius semicupreus
      • Chlaenius chomthongensis
      • Chlaenius glaucinus
      • Chlaenius raffrayi
      • Chlaenius noguchii
      • Chlaenius cribellicollis
      • A Ground Beetle (Chlaenius purpuricollis)
      • Chlaenius simplex
      • Chlaenius tiomanensis
      • Chlaenius manowianus
      • Chlaenius pallipes
      • Chlaenius analis
      • Chlaenius paulyi
      • Chlaenius saginatus
      • Chlaenius doriae
      • Chlaenius sivorii
      • Chlaenius oodioides
      • Chlaenius radama
      • Chlaenius sumptuosus
      • Chlaenius nitidifrons
      • Chlaenius gaoligongensis
      • Ground beetle (Chlaenius erythropus)
      • Chlaenius australis
      • Chlaenius ruvumaensis
      • Chlaenius darlingtoni
      • Chlaenius electrinus
      • Chlaenius merkli
      • Chlaenius tibialis
      • Chlaenius lastii
      • Chlaenius signatus
      • Chlaenius rampurensis
      • Chlaenius laetoides
      • Chlaenius controversus
      • Chlaenius montanus
      • Chlaenius cherensis
      • Chlaenius aestivus
      • Chlaenius langmusiensis
      • Chlaenius ivorensis
      • Chlaenius aurifex
      • Chlaenius morio
      • Chlaenius cham
      • Chlaenius montivagus
      • Chlaenius marginellus
      • Chlaenius diabolicus
      • Chlaenius boali
      • Chlaenius sibuti
      • Chlaenius rukwaensis
      • Chlaenius nigripennis
      • Chlaenius sankuruensis
      • Chlaenius xanthacrus
      • Chlaenius baeticus
      • Chlaenius howa
      • Chlaenius analisimilis
      • Chlaenius loxias
      • Chlaenius philemon
      • Chlaenius terminatus
      • Chlaenius schuelei
      • Chlaenius eggeri
      • Chlaenius forreri
      • Chlaenius rugulosus
      • Chlaenius marginicollis
      • Chlaenius maxi
      • Chlaenius manus
      • Chlaenius laeviplaga
      • Chlaenius indutus
      • Chlaenius rudesculptus
      • Chlaenius freynei
      • Chlaenius chapanus
      • Chlaenius wegneri
      • Chlaenius effugiens
      • Chlaenius jureceki
      • Chlaenius bengalensis
      • Chlaenius fairmairei
      • Chlaenius limbicollis
      • Chlaenius wrasei
      • Chlaenius pacholatkoi
      • Chlaenius kirschenhoferi
      • Chlaenius makokou
      • Chlaenius neocaledonicus
      • Chlaenius praefectus
      • Chlaenius columbinus
      • Chlaenius shaanxinensis
      • Chlaenius olthofi
      • Chlaenius crebrepunctatus
      • Chlaenius protensus
      • Chlaenius gambelaensis
      • Chlaenius sinensis
      • Chlaenius oculatus
      • Chlaenius yangonensis
      • Chlaenius jordani
      • Chlaenius armenus
      • Chlaenius acutecostatus
      • Chlaenius caeruleiceps
      • Chlaenius caurinus
      • Chlaenius heidenfelderi
      • Chlaenius davidsoni
      • Chlaenius celer
      • Chlaenius colasi
      • Chlaenius evae
      • Chlaenius brevior
      • Chlaenius gikondo
      • Chlaenius hildebrandti
      • Chlaenius ripicola
      • Chlaenius cursor
      • Chlaenius posticalis
      • Chlaenius victor
      • Chlaenius rambouseki
      • Chlaenius baoxingensis
      • Chlaenius suensoni
      • Chlaenius nanlingensis
      • Chlaenius wenchuanensis
      • Chlaenius pailin
      • Chlaenius suberbiei
      • Chlaenius caffer
      • Chlaenius fimbriatus
      • Chlaenius virens
      • Chlaenius atripes
      • Ground beetle (Chlaenius tricolor)
      • Chlaenius trigonotomoides
      • Chlaenius cupreolineatus
      • Chlaenius cookei
      • Chlaenius alluaudi
      • Chlaenius gabonicus
      • Chlaenius kerkvoordeae
      • Chlaenius goossensi
      • Chlaenius wallacei
      • Ground beetle (Chlaenius tomentosus)
      • Chlaenius maculatus
      • Chlaenius leishanensis
      • Chlaenius baliensis
      • Chlaenius waddellii
      • Chlaenius stungtrengensis
      • Chlaenius pyrrhopodus
      • Chlaenius luzonicus
      • Chlaenius sennaariensis
      • Chlaenius quadrinotatus
      • Chlaenius ochroperas
      • Chlaenius vitalisi
      • Chlaenius syriacus
      • Chlaenius boukali
      • Chlaenius kryzhanovskyi
      • Chlaenius breviusculus
      • Chlaenius geniculatus
      • Chlaenius taveuniensis
      • Chlaenius decorsei
      • Chlaenius vividus
      • Chlaenius cyaneus
      • Chlaenius trichrous
      • Chlaenius lulengae
      • Chlaenius judianensis
      • Chlaenius liothorax
      • Chlaenius prabangensis
      • Chlaenius loeblianus
      • Chlaenius zygogrammus
      • Chlaenius ernesti
      • Chlaenius fugax
      • Chlaenius augustus
      • Chlaenius gestroi
      • Chlaenius elgonensis
      • Chlaenius accedens
      • Chlaenius pongraczi
      • Chlaenius patrizii
      • Chlaenius floridanus
      • Chlaenius obsidianus
      • Chlaenius ditulus
      • Chlaenius rubenticollis
      • Chlaenius pseudolulengae
      • Chlaenius palpalis
      • Chlaenius perpunctatus
      • Chlaenius crudelis
      • Chlaenius luteoapicalis
      • Chlaenius boisduvalii
      • Chlaenius quadriornatus
      • Chlaenius tetragonoderus
      • Chlaenius mutatus
      • Chlaenius intermedius
      • Chlaenius chengduensis
      • Chlaenius kwazuluensis
      • Chlaenius junceus
      • Chlaenius perrieri
      • Chlaenius congoensis
      • Chlaenius systolocranioides
      • Chlaenius scheerpeltzi
      • Chlaenius opisthographus
      • Chlaenius tschitscherini
      • Chlaenius guatemalenus
      • Chlaenius transvaalensis
      • Chlaenius agilis
      • Chlaenius omochlorus
      • Chlaenius aterrimus
      • Chlaenius chiangmaiensis
      • Chlaenius lyperus
      • Chlaenius lirifer
      • Chlaenius nigrosuturatus
      • Chlaenius sexguttatus
      • Chlaenius puncticephalis
      • Chlaenius plicatipennis
      • Chlaenius basilimbatus
      • Chlaenius ocreatus
      • Chlaenius douei
      • Chlaenius amauropterus
      • Chlaenius alesi
      • Chlaenius lissoderus
      • Chlaenius veselyi
      • Chlaenius akanensis
      • Chlaenius sericimicans
      • Chlaenius savanicola
      • Chlaenius amplicollis
      • Chlaenius impressicollis
      • Chlaenius parallelus
      • Chlaenius brasiliensis
      • Chlaenius silvestrii
      • Ground beetle (Chlaenius emarginatus)
      • Chlaenius etoshaensis
      • Chlaenius corrosulus
      • Chlaenius fulgidicollis
      • Chlaenius pectinipes
      • Chlaenius wewalkai
      • Chlaenius caecus
      • Chlaenius dusaultii
      • Chlaenius olivieri
      • Chlaenius siccus
      • Ground beetle (Chlaenius pusillus)
      • Chlaenius densaticollis
      • Chlaenius comans
      • Chlaenius suvorovi
      • Chlaenius curtulus
      • Chlaenius tchabalensis
      • Chlaenius cosciniophorus
      • Chlaenius sculptilis
      • Chlaenius glabricollis
      • Chlaenius uninotatus
      • Chlaenius flavoscriptus
      • Chlaenius oligochrysus
      • Chlaenius pallidecornis
      • Chlaenius lunatus
      • Chlaenius tinantae
      • Chlaenius orbus
      • Chlaenius andamanensis
      • Chlaenius albissoni
      • Chlaenius chuanqianensis
      • Chlaenius puchneri
      • Chlaenius elongatulus
      • Chlaenius monardi
      • Chlaenius principalis
      • Chlaenius corbetti
      • Chlaenius leprieuri
      • Chlaenius freyellus
      • Chlaenius victoriae
      • Chlaenius gundlachi
      • Chlaenius melanopus
      • Chlaenius exaratus
      • Chlaenius laeticollis
      • Chlaenius semicyaneus
      • Chlaenius discopictus
      • Chlaenius kirki
      • Chlaenius consors
      • Chlaenius dissidens
      • Chlaenius viridicollis
      • Chlaenius laeviusculus
      • Chlaenius zanzibaricus
      • Chlaenius athleta
      • Chlaenius difficilis
      • Chlaenius validicornis
      • Chlaenius nepalensis
      • Chlaenius prasinus
      • Chlaenius kotys
      • Chlaenius pendjariensis
      • Chlaenius bakwuensis
      • Chlaenius attenuatus
      • Chlaenius leucoristus
      • Chlaenius pseudotristis
      • Chlaenius adametzi
      • Chlaenius ziloensis
      • Chlaenius fulvicollis
      • Chlaenius umtalianus
      • Chlaenius pseudobipustulatus
      • Chlaenius momboensis
      • Chlaenius probsti
      • Chlaenius rotundatulus
      • Chlaenius perinetanus
      • Chlaenius alutaceus
      • Chlaenius dimidiatus
      • Chlaenius stichai
      • Chlaenius germanus
      • Chlaenius vadoni
      • Chlaenius latipalpis
      • Chlaenius dilutipes
      • Chlaenius tshibindensis
      • Chlaenius linwensini
      • Chlaenius skukuzaensis
      • Chlaenius merklianus
      • Chlaenius venustulus
      • Chlaenius fouquei
      • Chlaenius braminus
      • Chlaenius jaleswarensis
      • Chlaenius tecospilus
      • Chlaenius luculentus
      • Chlaenius transfuga
      • Chlaenius sexmaculatus
      • Chlaenius klapperichi
      • Chlaenius allardi
      • Chlaenius semipurpureus
      • Chlaenius chalcoderus
      • Chlaenius aberrans
      • Chlaenius vertagoides
      • A Ground Beetle (Chlaenius harpalinus)
      • Chlaenius schmidti
      • Chlaenius straneoi
      • Chlaenius varians
      • Chlaenius rafiki
      • Chlaenius crenistriatus
      • Chlaenius quadrimaculatus
      • Chlaenius porrectus
      • Chlaenius maowensis
      • Chlaenius guineensis
      • Chlaenius brunnescens
      • Ground beetle (Chlaenius cordicollis)
      • Chlaenius leucops
      • Chlaenius dureli
      • Chlaenius reichei
      • Chlaenius privatus
      • Chlaenius festivus
      • Chlaenius differens
      • Chlaenius cheni
      • Chlaenius lugens
      • Chlaenius gonospilus
      • Chlaenius congoanus
      • Chlaenius multicolor
      • Chlaenius persimilis
      • Chlaenius specularis
      • Chlaenius masoni
      • A Ground Beetle (Chlaenius platyderus)
      • Chlaenius atratus
      • Chlaenius pumilio
      • Chlaenius lineatus
      • Chlaenius henryi
      • Chlaenius cordifer
      • Chlaenius vethi
      • Chlaenius auripilis
      • Chlaenius schillhammeri
      • Chlaenius katanga
      • Chlaenius bisignatus
      • Chlaenius tuky
      • Chlaenius infantulus
      • Chlaenius latreillei
      • Chlaenius lapillus
      • Chlaenius costiger
      • Chlaenius pulchellus
      • Chlaenius pseudoglabratus
      • Chlaenius cylindricollis
      • Chlaenius bohemani
      • Chlaenius fletcheri
      • Chlaenius greyanus
      • Chlaenius maxillosus
      • Chlaenius kolbei
      • Chlaenius thiesensis
      • Chlaenius eruditus
      • Chlaenius loveridgei
      • Chlaenius amydrus
      • Chlaenius oblongus
      • Chlaenius fraterculus
      • Chlaenius lucidulus
      • Chlaenius modestus
      • Chlaenius patricius
      • Chlaenius chrysocephalus
      • A Ground Beetle (Chlaenius interruptus)
      • Chlaenius sibutensis
      • Chlaenius amplipennis
      • Chlaenius kivuanus
      • Chlaenius atropos
      • Chlaenius latipennis
      • Chlaenius lioderus
      • Chlaenius trinotatus
      • Chlaenius afganus
      • Chlaenius obenbergeri
      • Chlaenius aodai
      • Chlaenius planulatus
      • Chlaenius betioky
      • Chlaenius dohrnii
      • Chlaenius daressalaami
      • Chlaenius lucidicollis
      • Chlaenius dichrous
      • Chlaenius tenuicollis
      • Chlaenius malcheri
      • Chlaenius pleuroderus
      • Chlaenius nebraskensis
      • Chlaenius coquerelii
      • Chlaenius orphanus
      • Chlaenius circumcinctus
      • Chlaenius cambodiensis
      • Chlaenius longeantennatus
      • Chlaenius morosus
      • Chlaenius simulatorius
      • Chlaenius assecla
      • Chlaenius colmanti
      • Chlaenius fizianus
      • Chlaenius pseudopoeciloides
      • Chlaenius obesus
      • Chlaenius uzungwensis
      • Chlaenius paykulli
      • Chlaenius treichi
      • Chlaenius leigongshanensis
      • Chlaenius helvetorum
      • Chlaenius alaotraensis
      • Chlaenius prostenus
      • Chlaenius meticulosus
      • Chlaenius pseudocruciatus
      • Chlaenius schoutedeni
      • Chlaenius panagaeoides
      • Chlaenius nitidulus
      • Chlaenius glaucus
      • Chlaenius maputoensis
      • Chlaenius cecrops
      • Chlaenius collarti
      • Chlaenius chrysoderus
      • Chlaenius suavis
      • Chlaenius himalayicus
      • Chlaenius scapularis
      • Chlaenius variolosus
      • Chlaenius mediornatus
      • Chlaenius variabilipes
      • Chlaenius startellus
      • Chlaenius rodriguezi
      • Chlaenius viduus
      • Chlaenius pertinax
      • Chlaenius sagaingensis
      • Chlaenius dives
      • Chlaenius nimbanus
      • Chlaenius nitidicollis
      • Chlaenius bocandei
      • Chlaenius timorensis
      • Chlaenius rhodesianus
      • Chlaenius bodhidharma
      • Chlaenius wittei
      • Chlaenius epistrophus
      • Chlaenius semenowi
      • Chlaenius exilis
      • Chlaenius infersus
      • Chlaenius lombokensis
      • Chlaenius aspericollis
      • Chlaenius obtusus
      • Chlaenius soricinus
      • Chlaenius sabahensis
      • Chlaenius aberanus
      • Chlaenius peruanus
      • Chlaenius samoensis
      • Chlaenius gunganagarensis
      • Chlaenius eryx
      • Chlaenius micans
      • Chlaenius notula
      • Chlaenius violatus
      • Chlaenius subsulcatus
      • Chlaenius yunnanulus
      • Chlaenius punctulatus
      • Chlaenius kira
      • Chlaenius kilimanus
      • Chlaenius formosensis
      • Chlaenius quadrisignatus
      • Chlaenius anischenkoi
      • Chlaenius pan
      • Chlaenius perspicillaris
      • Chlaenius delicatus
      • Chlaenius scotti
      • Chlaenius bilyi
      • Chlaenius clypeopatens
      • Chlaenius pseudopalpalis
      • Chlaenius mandli
      • Chlaenius superstes
      • Chlaenius puncticollis
      • Chlaenius medioguttatus
      • Ground beetle (Chlaenius nemoralis)
      • Chlaenius dostojevskii
      • Chlaenius chlorochrous
      • Chlaenius immaculatus
      • Chlaenius deplanatus
      • Chlaenius croesus
      • Chlaenius piceus
      • Chlaenius ghesquierei
      • Chlaenius betrokaensis
      • Chlaenius li
      • Chlaenius richardsi
      • Chlaenius inops
      • Chlaenius diversus
      • Chlaenius bangkokensis
      • Ground beetle (Chlaenius impunctifrons)
      • Chlaenius ovampo
      • Chlaenius planipennis
      • Chlaenius glabratus
      • Chlaenius malachinus
      • Chlaenius poecilinus
      • Chlaenius lateralis
      • Chlaenius mkongoensis
      • Chlaenius simbabwensis
      • Chlaenius propinquus
      • Chlaenius jucundulus
      • Chlaenius laetus
      • Chlaenius abacoides
      • Chlaenius benyovszkyi
      • Chlaenius dissimilis
      • Chlaenius hostilis
      • Chlaenius tamdaoensis
      • Chlaenius rudicollis
      • Chlaenius testaceicrus
      • Chlaenius canariensis
      • Chlaenius chuji
      • Chlaenius deyrollei
      • Chlaenius newdehliensis
      • Chlaenius garnerae
      • Chlaenius chainatensis
      • Chlaenius jacobsoni
      • Chlaenius costulatus
      • Chlaenius tenuelimbatus
      • Chlaenius posticus
      • Chlaenius latesternalis
      • Chlaenius koenigi
      • Chlaenius sinuatus
      • Chlaenius elongatus
      • Chlaenius sonomae
      • Chlaenius peltastes
      • Chlaenius kolariensis
      • Chlaenius robustus
      • Chlaenius monachus
      • Chlaenius quadripustulatus
      • Chlaenius pudicus
      • Chlaenius platynoides
      • Chlaenius subovatus
      • Chlaenius seiferti
      • Chlaenius sanatonialis
      • Chlaenius pulcher
      • Chlaenius djaina
      • Chlaenius sehnali
      • Chlaenius cavilabrum
      • Chlaenius comes
      • Chlaenius nilgiricus
      • Chlaenius leytensis
      • Chlaenius nigroscelis
      • Chlaenius hamifer
      • Chlaenius marleyi
      • Chlaenius lamottei
      • Chlaenius deliciolus
      • Chlaenius pterostichoides
      • Chlaenius imitatus
      • Chlaenius vitticollis
      • Chlaenius menevillei
      • Chlaenius punctatostriatus
      • Chlaenius siffointei
      • Chlaenius inderiensis
      • Chlaenius seyrigi
      • Chlaenius nubicus
      • Chlaenius deuvei
      • Chlaenius makalolo
      • Chlaenius pubipennis
      • Chlaenius orbicollis
      • Chlaenius assamensis
      • Chlaenius aeneocephalus
      • Chlaenius vietnami
      • Chlaenius robertae
      • Chlaenius ostrinus
      • Chlaenius curtii
      • Chlaenius dajuensis
      • Chlaenius trachys
      • Chlaenius incognitus
      • Chlaenius eugrammus
      • Chlaenius microspilus
      • Ground beetle (Chlaenius alternatus)
      • Chlaenius tsanerenaensis
      • Chlaenius convexus
      • Chlaenius chlorodius
      • Chlaenius iranensis
      • Chlaenius janthinus
      • Chlaenius flaccidus
      • Chlaenius hemichlorus
      • Chlaenius dispreticus
      • Chlaenius propeagilis
      • Chlaenius aethiopicus
      • Chlaenius jaegeri
      • Chlaenius noeli
      • Chlaenius cribricollis
      • Chlaenius toubaensis
      • Chlaenius sciakyi
      • Chlaenius tetraspilus
      • Chlaenius virgulatus
      • Chlaenius liocephalus
      • Chlaenius paluensis
      • Chlaenius eyeni
      • Chlaenius platensis
      • Chlaenius sokotranus
      • Chlaenius inaequalis
      • Chlaenius kurosawai
      • Chlaenius elegans
      • Chlaenius hypocrita
      • Chlaenius syangyaensis
      • Chlaenius piligenys
      • Chlaenius pilosicoerulans
      • Chlaenius extensus
      • Chlaenius nebrioides
      • Chlaenius amarae
      • Chlaenius milloti
      • Chlaenius prolixus
      • Chlaenius binghami
      • Chlaenius peterseni
      • Chlaenius kashmiricus
      • Chlaenius obsoletus
      • Chlaenius horni
      • Chlaenius boueti
      • Chlaenius martinbaehri
      • Chlaenius madecassus
      • Chlaenius trapezicollis
      • Chlaenius coeruleus
      • Chlaenius sagittarius
      • Chlaenius rotundithorax
      • Chlaenius dejeanii
      • Chlaenius variicornis
      • Chlaenius rufifemoratus
      • Chlaenius splendidus
      • Chlaenius angolanus
      • Chlaenius soginoides
      • Chlaenius spoliatus
      • Chlaenius patruelis
      • Chlaenius hainanensis
      • Chlaenius euryscopus
      • Chlaenius isaloensis
      • Chlaenius yamdena
      • Chlaenius retropictus
      • Chlaenius kanarae
      • Chlaenius feronoides
      • Chlaenius hamatus
      • Chlaenius bottegoi
      • Chlaenius kivuensis
      • Chlaenius nigratus
      • Chlaenius bergerdostali
      • Chlaenius alexanderdostali
      • Chlaenius senegalensis
      • Chlaenius postmaculatus
      • Chlaenius limbatus
      • Chlaenius laevicollis
      • Chlaenius lineellus
      • Chlaenius meyeri
      • Chlaenius jakeschi
      • Chlaenius orbiculicollis
      • Chlaenius toliaraensis
      • Chlaenius gemmingeri
      • Chlaenius backanensis
      • Chlaenius moaboensis
      • Chlaenius udaipurensis
      • Chlaenius nigrans
      • Chlaenius fastigatus
      • Chlaenius borellyi
      • Chlaenius janaki
      • Chlaenius synaptus
      • Chlaenius hunanensis
      • Chlaenius jaechi
      • Chlaenius pandagensis
      • Chlaenius conformis
      • Chlaenius sollicitus
      • Chlaenius chaudoiri
      • Chlaenius madangensis
      • Chlaenius cruralis
      • Chlaenius coxalis
      • Chlaenius touzalini
      • Chlaenius cupreolus
      • Chlaenius phenax
      • Chlaenius milleti
      • Chlaenius eritreaensis
      • Chlaenius naeviger
      • Chlaenius guttula
      • Chlaenius cyanostolus
      • Chlaenius antennatus
      • Chlaenius ptuchodes
      • Ground beetle (Chlaenius laticollis)
      • Chlaenius pratensis
      • Chlaenius laotinus
      • Chlaenius quadricolor
      • Chlaenius viridis
      • Chlaenius marianensis
      • Chlaenius bandjermasinensis
      • Chlaenius cupreocinctus
      • Chlaenius leptopus
      • Chlaenius plausibilis
      • Chlaenius goryi
      • Chlaenius caesitius
      • Chlaenius humphreyi
      • Chlaenius gabonensis
      • Chlaenius camerunensis
      • Chlaenius overlaeti
      • Chlaenius instabilis
      • Chlaenius distigma
      • Chlaenius daer
      • Chlaenius opacipennis
      • Chlaenius kafakumbae
      • Chlaenius anthracoderus
      • Chlaenius sulcicollis
      • Chlaenius semperi
      • Chlaenius sallei
      • Chlaenius consobrinus
      • Chlaenius qinqchensis
      • Chlaenius contractus
      • Chlaenius melanopterus
      • Chlaenius latithorax
      • Chlaenius louwerensi
      • Chlaenius kenyerii
      • Chlaenius bipustulatus
      • Chlaenius communimacula
      • Ground beetle (Chlaenius niger)
      • Chlaenius fulvipes
      • Chlaenius kaszabi
      • Chlaenius argentinicus
      • Chlaenius aruwimius
      • Chlaenius stevenii
      • Chlaenius insulanus
      • Chlaenius conradsi
      • Chlaenius lacunosus
      • Chlaenius quadrivittatus
      • Chlaenius kulti
      • Chlaenius farai
      • Chlaenius greensladei
      • Chlaenius erythrocnemis
      • Chlaenius bulirschi
      • Chlaenius togatus
      • Chlaenius pyrrhos
      • Chlaenius watsaensis
      • Chlaenius ruficauda
      • Chlaenius elisabethanus
      • Chlaenius swahilius
      • Chlaenius ranavalonae
      • Chlaenius titschacki
      • Chlaenius reitteri
      • Chlaenius lujai
      • Chlaenius marondera
      • A Ground Beetle (Chlaenius solitarius)
      • Chlaenius moluccensis
      • Chlaenius bivulnerus
      • Chlaenius kathrynae
      • Chlaenius amazonicus
      • Chlaenius herbaceus
      • Chlaenius simillimus
      • Chlaenius lucasii
      • Chlaenius velutinus
      • Chlaenius vulneratus
      • Chlaenius jeanneli
      • Chlaenius championi
      • Chlaenius fenestratus
      • Chlaenius geisthardti
      • Chlaenius ceramensis
      • Chlaenius arnoldi
      • Chlaenius rapyllii
      • Chlaenius allardianus
      • Chlaenius meteorus
      • Chlaenius melampus
      • Chlaenius angustatus
      • Chlaenius impictus
      • Chlaenius dilatatus
      • Chlaenius kathmanduensis
      • Chlaenius procerulus
      • Chlaenius pericallus
      • Chlaenius katanganus
      • Chlaenius somaliae
      • Chlaenius fasciger
      • Chlaenius dorsalis
      • Chlaenius pachys
      • Chlaenius flavomaculatus
      • Chlaenius fraternus
      • Chlaenius eurybates
      • Chlaenius drescheri
      • Chlaenius cribellatus
      • Chlaenius berndjaegeri
      • Chlaenius postscriptus
      • Chlaenius annulipes
      • Chlaenius adagidensis
      • Chlaenius beatus
      • Chlaenius cebuensis
    • Callistus lunatus
  • Licinini → (178 species)
    • Badister → (56 species)
      • Badister pictus
      • Badister maculatus
      • Badister marginellus
      • Badister dorsiger
      • Ground beetle (Badister micans)
      • Badister prodromus
      • Badister cavifrons
      • Badister iridescens
      • Badister naviauxi
      • Badister unipustulatus
      • Badister ishigakiensis
      • Badister antecursor
      • Ground beetle (Badister pulchellus)
      • Badister sasajii
      • Badister nigriceps
      • Badister gladiator
      • Badister debilis
      • Badister thoracicus
      • Badister mareei
      • Badister macrocephalus
      • Badister collaris
      • Ground beetle (Badister neopulchellus)
      • Badister elegans
      • Badister vittatus
      • Badister sundaicus
      • Badister dilatatus
      • Badister nakayamai
      • Badister grandis
      • Badister bucciarellii
      • Badister flavipes
      • Badister seriepunctatus
      • Badister ajax
      • Badister amazonus
      • Ground beetle (Badister obtusus)
      • Badister fenestratus
      • Badister bullatus
      • Badister lacertosus
      • A Ground Beetle (Badister transversus)
      • Ground beetle (Badister reflexus)
      • A Ground Beetle (Badister ferrugineus)
      • Badister anatolicus
      • Badister promontorii
      • Badister iranicus
      • Badister brevicollis
      • Badister fragilis
      • Badister denticulatus
      • Ground beetle (Badister grandiceps)
      • Badister peltatus
      • Badister ussuriensis
      • Badister fukiensis
      • Badister sodalis
      • Badister vandykei
      • Ground beetle (Badister parviceps)
      • Badister meridionalis
      • Badister submarinus
      • Ground beetle (Badister notatus)
    • Lestignathus → (4 species)
      • Lestignathus pieperi
      • Lestignathus foveatus
      • Lestignathus cursor
      • Lestignathus simsoni
    • Lacordairia → (12 species)
      • Lacordairia taylori
      • Lacordairia erichsoni
      • Lacordairia elongata
      • Lacordairia anchomenoides
      • Lacordairia fugax
      • Lacordairia calathoides
      • Lacordairia terrena
      • Lacordairia angustata
      • Lacordairia proxima
      • Lacordairia cychroides
      • Lacordairia insulicola
      • Lacordairia argutoroides
    • Diplocheila → (28 species)
      • Diplocheila perscissa
      • Diplocheila polita
      • Diplocheila obtusa
      • A Ground Beetle (Diplocheila oregona)
      • Diplocheila cordicollis
      • A Ground Beetle (Diplocheila undulata)
      • Diplocheila latifrons
      • Diplocheila laevis
      • A Ground Beetle (Diplocheila striatopunctata)
      • Diplocheila pinodes
      • Diplocheila henshawi
      • Diplocheila aegyptiaca
      • Diplocheila zealandica
      • Diplocheila exotica
      • Diplocheila minima
      • Ground beetle (Diplocheila major)
      • Diplocheila laevigata
      • Diplocheila crossi
      • Diplocheila colossus
      • Diplocheila transcaspica
      • A Ground Beetle (Diplocheila impressicollis)
      • Diplocheila nupera
      • Diplocheila capensis
      • Diplocheila laevigotoides
      • Diplocheila elongata
      • Diplocheila zeelandica
      • Diplocheila distinguenda
      • Diplocheila macromandibularis
    • Dicaelus → (18 species)
      • Dicaelus alternans
      • Dicaelus crenatus
      • Dicaelus chermocki
      • A Ground Beetle (Dicaelus purpuratus)
      • A Ground Beetle (Dicaelus laevipennis)
      • Dicaelus quadratus
      • Dicaelus franclemonti
      • Dicaelus dilatatus
      • Dicaelus alutaceus
      • Ground beetle (Dicaelus elongatus)
      • Ground beetle (Dicaelus politus)
      • A Ground Beetle (Dicaelus sculptilis)
      • A Ground Beetle (Dicaelus teter)
      • Dicaelus costatus
      • Dicaelus subtropicus
      • Dicaelus suffusus
      • Dicaelus ambiguus
      • Dicaelus furvus
    • Dicrochile → (30 species)
      • Dicrochile novaezelandiae
      • Dicrochile tiro
      • Dicrochile subopaca
      • Dicrochile cordicollis
      • Dicrochile gigas
      • Dicrochile cephalotes
      • Dicrochile minuta
      • Dicrochile maura
      • Dicrochile idae
      • Dicrochile aterrima
      • Dicrochile artensis
      • Dicrochile anthracina
      • Dicrochile whitei
      • Dicrochile punctatostriata
      • Dicrochile brevicollis
      • Dicrochile insignis
      • Dicrochile alternans
      • Dicrochile goryi
      • Dicrochile punctipennis
      • Dicrochile fabrii
      • Dicrochile ventralis
      • Dicrochile quadricollis
      • Dicrochile nitida
      • Dicrochile flavipes
      • Dicrochile thoracica
      • Dicrochile rugicollis
      • Dicrochile punctulata
      • Dicrochile acuta
      • Dicrochile anchomenoides
      • Dicrochile caledonica
    • Licinus → (29 species)
      • Licinus silphoides
      • Licinus oertzeni
      • Licinus bartoni
      • Licinus punctatulus
      • Licinus graecus
      • Licinus merkli
      • Licinus afghanistanus
      • Licinus depressus
      • Licinus manriquianus
      • Licinus cassideus
      • Licinus corustes
      • Licinus italicus
      • Licinus mongolicus
      • Licinus iranicus
      • Licinus planicollis
      • Licinus aequatus
      • Licinus astrabadensis
      • Licinus setosus
      • Licinus yezoensis
      • Licinus oblongus
      • Licinus hoffmannseggii
      • Licinus cordatus
      • Licinus schuberti
      • Licinus lindbergi
      • Licinus jaloricus
      • Licinus convexus
      • Licinus pubifer
      • Licinus aegyptiacus
      • Licinus peltoides
    • Eutogeneius fuscus
  • Oodini → (130 species)
    • Dercylus → (35 species)
      • Dercylus praepilatus
      • Dercylus chaudoiri
      • Dercylus gibber
      • Dercylus tenebricosus
      • Dercylus alternans
      • Dercylus convexus
      • Dercylus catenatus
      • Dercylus infernus
      • Dercylus mathani
      • Dercylus onorei
      • Dercylus gaujoni
      • Dercylus ohausi
      • Dercylus davidsoni
      • Dercylus franiai
      • Dercylus batesii
      • Dercylus gibbosus
      • Dercylus granifer
      • Dercylus cordicollis
      • Dercylus italoi
      • Dercylus steinbachi
      • Dercylus punctatosulcatus
      • Dercylus mexicanus
      • Dercylus bolivianus
      • Dercylus opacus
      • Dercylus puritanus
      • Dercylus crenatus
      • Dercylus orbiculatus
      • Dercylus nodosus
      • Dercylus licinoides
      • Dercylus tuberculatus
      • Dercylus heynei
      • Dercylus buckleyi
      • Dercylus ater
      • Dercylus anthracinus
      • Dercylus gautardi
    • Stenocrepis → (30 species)
      • Stenocrepis pauper
      • Stenocrepis olivacea
      • Stenocrepis angustipennis
      • Stenocrepis insulana
      • Stenocrepis gratiosa
      • Stenocrepis subdepressa
      • Stenocrepis aeruginea
      • Stenocrepis elegans
      • Stenocrepis laevigata
      • Stenocrepis palustris
      • Stenocrepis guerini
      • Stenocrepis sahlbergii
      • Stenocrepis viridula
      • Stenocrepis pallipes
      • Stenocrepis leprieurii
      • Stenocrepis gilvipes
      • Stenocrepis punctatostriata
      • Stenocrepis cayennensis
      • Stenocrepis nigricornis
      • Stenocrepis fuscipes
      • Stenocrepis robusta
      • Stenocrepis marginella
      • Stenocrepis sinuata
      • Stenocrepis metallica
      • Stenocrepis triaria
      • Stenocrepis flavicrus
      • Stenocrepis mexicana
      • Stenocrepis duodecimstriata
      • Stenocrepis quatuordecimsulcata
      • Stenocrepis cuprea
    • Oodes → (50 species)
      • Oodes angustus
      • Oodes siccus
      • Oodes congoensis
      • Oodes desertus
      • Oodes wilsoni
      • Oodes angustatus
      • Oodes trisulcatus
      • Ground beetle (Oodes americanus)
      • Oodes calvus
      • Oodes sulcicollis
      • Oodes oblongus
      • Oodes helopioides
      • Oodes longus
      • Oodes modestus
      • Oodes xanthochilus
      • Oodes taprobanae
      • Oodes walshae
      • Oodes lenis
      • Oodes irakensis
      • Oodes nil
      • Ground beetle (Oodes brevis)
      • Oodes inornatus
      • Ground beetle (Oodes fluvialis)
      • Oodes paroensis
      • Oodes palpalis
      • Oodes coelestinus
      • Oodes bivittatus
      • Ground beetle (Oodes amaroides)
      • Oodes waterhousei
      • Oodes caerulans
      • Oodes subcoriaceus
      • Oodes cribristernis
      • Oodes parviceps
      • Oodes froggatti
      • Oodes vicarius
      • Oodes basrensis
      • Oodes melanodes
      • Oodes monticola
      • Oodes fitzroyensis
      • Oodes par
      • Oodes austrinus
      • Oodes denisonensis
      • Oodes terrestris
      • Oodes gracilis
      • Oodes rossi
      • Oodes bostockii
      • Oodes latior
      • Oodes sikkimensis
      • Oodes impressus
      • Oodes echigonus
    • Lachnocrepis → (3 species)
      • Lachnocrepis prolixa
      • Lachnocrepis parallela
      • Lachnocrepis japonica
    • Anatrichis → (11 species)
      • Anatrichis oblonga
      • Anatrichis morii
      • Anatrichis birmanica
      • Anatrichis indica
      • Anatrichis pedinoides
      • Anatrichis pusilla
      • Anatrichis minuta
      • Anatrichis nigra
      • Anatrichis infima
      • Anatrichis longula
      • Anatrichis ryukyuensis
    • Adelopomorpha glabra

Other links

• Wikidata
• Wikipedia (language: en)
• Wikipedia (language: fr)
• Wikipedia (language: nl)
• Encyclopedia of Life
• NCBI (genetics)"
• Open Tree of Life

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