Base of the evolutionary tree for Platyninae

← parent

• Platyninae (2094 species)
  • Platynini → (1375 species)
    • Platynus → (186 species)
      • Platynus macer
      • Platynus scriptellus
      • Platynus vagepunctatus
      • Platynus constricticeps
      • Platynus convexulus
      • Platynus biramosus
      • Platynus fractilinea
      • A Ground Beetle (Platynus ovipennis)
      • Platynus franiai
      • Platynus daviesi
      • Platynus ferghanicus
      • Platynus interitus
      • Platynus scriptus
      • Platynus calvini
      • Platynus depressus
      • Platynus trifoveolatus
      • Platynus klausnitzeri
      • Platynus amone
      • Platynus dianus
      • Platynus kleebergi
      • Platynus conicicollis
      • Platynus wassulandi
      • Platynus isthmiacus
      • Platynus schnitteri
      • Platynus transcibao
      • A Beetle (Platynus tenuicollis)
      • Platynus acuniai
      • Platynus cyanodorsalis
      • Platynus benardi
      • Platynus subovalis
      • Platynus kucerai
      • Platynus rubrofemoratus
      • Platynus krynickii
      • Platynus herculeanus
      • Platynus leiroides
      • Platynus deuvei
      • Platynus cohni
      • Platynus jamaicae
      • Platynus grassator
      • Platynus protensus
      • Platynus medius
      • Platynus complanatus
      • Platynus andrewesi
      • Platynus satsunanus
      • Platynus macropterus
      • Platynus jaegeri
      • Platynus parmarginatus
      • Platynus turberensis
      • Platynus arboreus
      • Platynus ramoni
      • Platynus senex
      • Platynus bruneri
      • Ground beetle (Platynus decentis)
      • Platynus klickai
      • Platynus florissantensis
      • Platynus azbleotroades
      • Platynus proximus
      • A Ground Beetle (Platynus brunneomarginatus)
      • Platynus halli
      • Platynus baragua
      • Platynus parallelosomus
      • Platynus punctus
      • Platynus dissipatus
      • Ground beetle (Platynus mannerheimii)
      • Platynus altifluminis
      • Platynus tartareus
      • Platynus aeneicauda
      • Platynus casus
      • Platynus rarus
      • Platynus angustatus
      • Platynus aeneosetosus
      • Platynus batesi
      • Platynus anatolicus
      • Platynus darlingtoni
      • Platynus opaculus
      • Platynus longaevus
      • Platynus scrobiculatus
      • Platynus zengae
      • Platynus kazuyoshii
      • Platynus tipoto
      • Platynus cychrinus
      • Platynus stricticollis
      • Platynus cinchonae
      • Platynus staveni
      • Platynus indecentis
      • Platynus takabai
      • Platynus fractilateralis
      • Platynus haemorrhous
      • Platynus pakistanensis
      • Platynus fratrorum
      • Platynus marcus
      • Platynus baroni
      • Platynus interglacialis
      • Platynus laeviceps
      • Platynus willbergi
      • Platynus caesus
      • Platynus pinarensis
      • Platynus subcordens
      • Platynus setiporus
      • Platynus teriolensis
      • Platynus livens
      • Platynus nuceus
      • Platynus incisus
      • Platynus subgelidus
      • Platynus lewisi
      • A Ground Beetle (Platynus cincticollis)
      • Platynus dilatipes
      • Platynus aguadensis
      • Platynus meurguesianus
      • Platynus robustus
      • Platynus praedator
      • Platynus tropicus
      • Platynus viridinitens
      • Platynus trisetosus
      • Platynus bromeliarum
      • Platynus montezumae
      • Platynus tolucensis
      • Platynus consularis
      • Platynus strictinotum
      • Platynus umbripennis
      • Platynus eupunctatus
      • Platynus harttii
      • Platynus komarovi
      • Platynus grandicollis
      • Platynus barclayi
      • Platynus prognathus
      • Platynus pygmaeus
      • Platynus cavicola
      • Platynus peirolerii
      • Platynus magnus
      • Platynus districtus
      • Platynus pecki
      • Platynus inops
      • Platynus ustus
      • Platynus pleistocenicus
      • Platynus cavatus
      • Platynus calathinus
      • Platynus exterminatus
      • Platynus carabiai
      • Platynus lyratus
      • Platynus ballorum
      • Platynus amicorum
      • Platynus viator
      • Platynus hamatus
      • Platynus visitor
      • Platynus bruskelchus
      • Platynus bucheri
      • Platynus mediopterus
      • Platynus lindrothi
      • Platynus subovatus
      • Platynus wolla
      • Platynus asper
      • Platynus hindei
      • Platynus insculptipennis
      • Platynus subangustus
      • Platynus desuetus
      • Platynus fur
      • Platynus henvelus
      • Platynus fukiensis
      • Platynus roysi
      • Platynus santarosae
      • Platynus dilapidatus
      • Platynus christophe
      • Platynus rastafarius
      • Platynus glacialis
      • Platynus laevis
      • Platynus sexualis
      • A Ground Beetle (Platynus hypolithos)
      • Platynus logicus
      • Platynus bruesi
      • Platynus acroglyptus
      • Platynus eulabes
      • Platynus cubensis
      • Platynus mixius
      • Platynus agilis
      • Platynus platynidioides
      • Platynus meurguesae
      • Platynus turquinensis
      • Platynus matsumurai
      • Platynus algirinus
      • Platynus richteri
      • Platynus tasmantus
      • Platynus lineopunctatus
      • Platynus faber
      • Platynus immarginatus
      • Platynus rougemonti
    • Ctenognathus → (32 species)
      • Ctenognathus bidens
      • Ctenognathus adamsi
      • Ctenognathus colensonis
      • Ctenognathus crenatus
      • Ctenognathus cheesemani
      • Ctenognathus novaezelandiae
      • Ctenognathus cardiophorus
      • Ctenognathus libitus
      • Ctenognathus littorellus
      • Ctenognathus sulcitarsis
      • Ctenognathus helmsi
      • Ctenognathus simmondsi
      • Ctenognathus edwardsii
      • Ctenognathus integratus
      • Ctenognathus otagoensis
      • Ctenognathus actochares
      • Ctenognathus intermedius
      • Ctenognathus parabilis
      • Ctenognathus lucifugus
      • Ctenognathus sophronitis
      • Ctenognathus politulus
      • Ctenognathus xanthomelas
      • Ctenognathus pictonensis
      • Ctenognathus deformipes
      • Ctenognathus arnaudensis
      • Ctenognathus suborbithorax
      • Ctenognathus sandageri
      • Ctenognathus perrugithorax
      • Ctenognathus neozelandicus
      • Ctenognathus macrocoelis
      • Ctenognathus oreobius
      • Ctenognathus munroi
    • Dicranoncus → (14 species)
      • Dicranoncus ravus
      • Dicranoncus quadridens
      • Dicranoncus philippinensis
      • Dicranoncus discoideus
      • Dicranoncus resplendens
      • Dicranoncus femoralis
      • Dicranoncus nigriventris
      • Dicranoncus pocillator
      • Dicranoncus bicolor
      • Dicranoncus atripennis
      • Dicranoncus vulpinus
      • Dicranoncus queenslandicus
      • Dicranoncus ganodes
      • Dicranoncus cinctipennis
    • Agonum → (204 species)
      • Agonum speculator
      • Agonum balesi
      • Agonum madagascariense
      • Agonum gracile
      • Agonum gisellae
      • Agonum quadripustulatum
      • Ground beetle (Agonum consimile)
      • Ground beetle (Agonum trigeminum)
      • Agonum persecretum
      • Ground beetle (Agonum thoreyi)
      • Agonum triimpressum
      • Agonum rigidulum
      • Agonum piceum
      • Agonum chalconotum
      • Agonum ericeti
      • Agonum bilimeki
      • Agonum luteum
      • Agonum shirahatai
      • Ground beetle (Agonum excavatum)
      • Agonum canadense
      • Agonum comatum
      • Agonum arisanum
      • Agonum rugicolle
      • Agonum charillus
      • Ground beetle (Agonum nutans)
      • Agonum chinense
      • Agonum nigrum
      • Ground beetle (Agonum striatopunctatum)
      • Agonum cheni
      • Agonum crenulatum
      • Ground beetle (Agonum metallescens)
      • Agonum yamatonis
      • Agonum sordidum
      • Agonum extimum
      • Agonum leucomerum
      • Agonum viridicupreum
      • Ground beetle (Agonum melanarium)
      • Agonum viridicans
      • A Ground Beetle (Agonum exaratum)
      • Agonum cyanopis
      • Agonum modestius
      • Ground beetle (Agonum fidele)
      • Agonum pacificum
      • Agonum jurecekianum
      • Agonum dolens
      • Ground beetle (Agonum affine)
      • Agonum antiquum
      • Agonum shibatai
      • Agonum viduum
      • Agonum carbonarium
      • Agonum hiranoi
      • Agonum sylphides
      • Agonum curtipenne
      • Ground beetle (Agonum tenue)
      • Agonum fallax
      • Agonum basale
      • six-point ground beetle (Agonum sexpunctatum)
      • Agonum liebherri
      • Agonum abnormale
      • Agonum deuvei
      • Agonum atlantis
      • Agonum aurelius
      • Ground beetle (Agonum palustre)
      • Agonum rugaticolle
      • Beller's Ground Beetle (Agonum belleri)
      • Agonum uenoi
      • Agonum fuliginosum
      • Agonum iriomotense
      • Agonum mandli
      • Agonum patinale
      • Agonum scutiferum
      • Agonum ferruginosum
      • Agonum azumai
      • Agonum parextimum
      • Agonum cyclifer
      • Agonum scitulum
      • Agonum rufipes
      • Agonum gerdmuelleri
      • Agonum sulcipenne
      • Ground beetle (Agonum errans)
      • Ground beetle (Agonum placidum)
      • Agonum chinese
      • Agonum obesum
      • Agonum scutifer
      • Ground beetle (Agonum retractum)
      • Agonum bulgani
      • Agonum guangdongense
      • Agonum maruokai
      • A Ground Beetle (Agonum picicornoides)
      • Agonum nipponicum
      • Agonum propinquum
      • Agonum deuveiellum
      • Agonum piceolum
      • Agonum hypocrita
      • Agonum emarginatum
      • Agonum scintillans
      • A Ground Beetle (Agonum deceptivum)
      • Ground beetle (Agonum muelleri)
      • Agonum menetriesii
      • Agonum kucerai
      • Agonum antennarium
      • Agonum praetor
      • Agonum illocatum
      • A Ground Beetle (Agonum mutatum)
      • Agonum chalcomum
      • Agonum mongolicum
      • Agonum gracilipes
      • Agonum versutum
      • Yellow-sided Marginatum (Agonum marginatum)
      • Agonum eurydamas
      • Agonum kurosonense
      • Agonum daimio
      • Agonum dreuxi
      • Ground beetle (Agonum corvus)
      • Agonum causticum
      • Ground beetle (Agonum albicrus)
      • Agonum texanum
      • Agonum lanei
      • Agonum apex
      • Agonum harrisii
      • A Ground Beetle (Agonum superioris)
      • Ground beetle (Agonum lutulentum)
      • Agonum limbatum
      • Ground beetle (Agonum nigriceps)
      • Agonum rufotestaceum
      • Ground beetle (Agonum sordens)
      • Agonum jankowskii
      • Agonum hirashimai
      • Agonum holzschuhi
      • Agonum platyderum
      • Agonum scotti
      • Agonum sylphis
      • Ground beetle (Agonum octopunctatum)
      • Agonum inaequale
      • Agonum lampros
      • Agonum sculptipes
      • Agonum aeruginosum
      • Agonum luczoti
      • Agonum triseriatum
      • Agonum pangxiongfeii
      • Agonum euroum
      • A Ground Beetle (Agonum moerens)
      • A Ground Beetle (Agonum brevicolle)
      • Agonum indiae
      • Agonum lanellum
      • Ground beetle (Agonum anchomenoides)
      • Agonum duftschmidi
      • Agonum ardoisi
      • Agonum mokoto
      • Agonum rougerieanum
      • Agonum suturale
      • Agonum nanum
      • A Ground Beetle (Agonum decorum)
      • Agonum dorsostriatum
      • Agonum humerosum
      • Agonum permoestum
      • Agonum sataense
      • Ground beetle (Agonum galvestonicum)
      • Agonum subtruncatum
      • Agonum rotundicolle
      • Agonum ishidai
      • Agonum amplior
      • A Ground Beetle (Agonum cupreum)
      • Agonum suavissimum
      • Agonum mesostictum
      • Agonum bicolor
      • Agonum japonicum
      • Agonum munsteri
      • Agonum kesharishresthae
      • Agonum semicupreum
      • Agonum imitans
      • Agonum anthracinum
      • Agonum tsushimanum
      • Agonum stenuiti
      • A Ground Beetle (Agonum darlingtoni)
      • Agonum micans
      • Agonum graciloides
      • Ground beetle (Agonum gratiosum)
      • Agonum tokarae
      • Agonum cupripenne
      • A Ground Beetle (Agonum crenistriatum)
      • Agonum deplanatum
      • Agonum quadrimaculatum
      • Agonum bellicum
      • Agonum afrum
      • Agonum atricomes
      • Agonum fossiger
      • A Ground Beetle (Agonum punctiforme)
      • Agonum teruyai
      • Agonum oryctus
      • Agonum angulifer
      • Agonum straeleni
      • Agonum numidicum
      • Ground beetle (Agonum extensicolle)
      • Ground beetle (Agonum ferreum)
      • Agonum shibataianum
      • Agonum lugens
      • Agonum collare
      • Agonum muiri
      • Agonum monachum
      • Agonum aequatum
      • A Ground Beetle (Agonum simile)
      • Agonum ladakense
      • Agonum longicorne
    • Limodromus → (3 species)
      • Limodromus longiventris
      • Limodromus hoffeinsorum
      • Limodromus assimilis
    • Paranchus → (4 species)
      • A Ground Beetle (Paranchus albipes)
      • Paranchus euthemon
      • Paranchus nichollsii
      • Paranchus debilis
    • Oxypselaphus → (4 species)
      • Oxypselaphus obscurus
      • Oxypselaphus dominici
      • A Ground Beetle (Oxypselaphus pusillus)
      • Oxypselaphus thielei
    • Notagonum → (80 species)
      • Notagonum fuscipes
      • Notagonum angulum
      • Notagonum garainae
      • Notagonum angustellum
      • Notagonum weigeli
      • Notagonum macleayi
      • Notagonum paludum
      • Notagonum marginellum
      • Notagonum margaritum
      • Notagonum ambulator
      • Notagonum darlingtoni
      • Notagonum piceum
      • Notagonum altum
      • Notagonum sigi
      • Notagonum iridius
      • Notagonum devosi
      • Notagonum excisipenne
      • Notagonum aitape
      • Notagonum skalei
      • Notagonum subangulum
      • Notagonum ullrichi
      • Notagonum luzonense
      • Notagonum subspinulum
      • Notagonum oxypterum
      • Notagonum reversior
      • Notagonum nigrinum
      • Notagonum subpunctum
      • Notagonum schuelei
      • Notagonum vile
      • Notagonum reversum
      • Notagonum macrops
      • Notagonum quadruum
      • Notagonum lawsoni
      • Notagonum mucidum
      • Notagonum subrufum
      • Notagonum lackneri
      • Notagonum vaporum
      • Notagonum gorokae
      • Notagonum subimpressum
      • Notagonum undatum
      • Notagonum anceps
      • Notagonum anops
      • submetallic ground beetle (Notagonum submetallicum)
      • Notagonum albertisi
      • Notagonum lafertei
      • Notagonum novaeguineae
      • Notagonum spinulum
      • Notagonum circumdatum
      • Notagonum astrum
      • Notagonum ilagae
      • Notagonum externum
      • Notagonum moluccense
      • Notagonum sinuum
      • Notagonum chathamense
      • Notagonum dentellum
      • Notagonum pereus
      • Notagonum gibbum
      • Notagonum curiosum
      • Notagonum laticolle
      • Notagonum nigrellum
      • Notagonum delaruei
      • Notagonum sectum
      • Notagonum feredayi
      • Notagonum laglaizei
      • Notagonum caritum
      • Notagonum inerme
      • Notagonum hamatum
      • Notagonum marginale
      • Notagonum pernitidum
      • Notagonum parvicolle
      • Notagonum malkini
      • Notagonum pleurale
      • Notagonum addendum
      • Notagonum drescheri
      • Notagonum exactum
      • Notagonum crenulipenne
      • Notagonum kitchingi
      • Notagonum subnigrum
      • Notagonum murrayense
      • Notagonum macrophthalmum
    • Calathidius → (4 species)
      • Calathidius accuminatus
      • Calathidius brevithorax
      • Calathidius acuminatus
      • Calathidius sphodroides
    • Glyptolenus → (34 species)
      • Glyptolenoides siemeradskii
      • Glyptolenoides cyclothorax
      • Glyptolenoides azureipennis
      • Glyptolenoides elegantulus
      • Glyptolenoides sulcitarsis
      • Glyptolenoides balli
      • Glyptolenoides azureus
      • Glyptolenoides germaini
      • Glyptolenoides purpuripennis
      • Glyptolenus transformatus
      • Glyptolenus affinis
      • Glyptolenus rivalis
      • Glyptolenus latitarsis
      • Glyptolenus smithi
      • Glyptolenus negrei
      • Glyptolenus apicestriatus
      • Glyptolenus janthinus
      • Glyptolenus chalybeus
      • Glyptolenus aereipennis
      • Glyptolenus spinosus
      • Glyptolenus chalybaeus
      • Glyptolenus rugicollis
      • Glyptolenus sulcipennis
      • Glyptolenus simplicicollis
      • Glyptolenus convexiusculus
      • Glyptolenus brevitarsis
      • Glyptolenus nigrita
      • Glyptolenus estebanensis
      • Glyptolenus nitidipennis
      • Glyptolenus ater
      • Glyptolenus ruficollis
      • Glyptolenus mirabilis
      • Glyptolenus latelytra
      • Glyptolenus straneoi
    • Dyscolus → (445 species)
      • Dyscolus anthracinus
      • Dyscolus lactipes
      • Dyscolus harpaloides
      • Dyscolus amplicollis
      • Dyscolus ellipticus
      • Dyscolus thecarum
      • Dyscolus cupripennis
      • Dyscolus longipes
      • Dyscolus niger
      • Dyscolus punctatostriatus
      • Dyscolus clarus
      • Dyscolus laevipennis
      • Dyscolus pavens
      • Dyscolus longipennis
      • Dyscolus sphodroides
      • Dyscolus tuberosus
      • Dyscolus cyanipennis
      • Dyscolus dyschirioides
      • Dyscolus rubellus
      • Dyscolus cyaneocupreus
      • Dyscolus leucoscelis
      • Dyscolus metallicus
      • Dyscolus costaricensis
      • Dyscolus aurotinctus
      • Dyscolus purulensis
      • Dyscolus bacatellus
      • Dyscolus pseudellipticus
      • Dyscolus duplex
      • Dyscolus brevis
      • Dyscolus mimulus
      • Dyscolus bracteatus
      • Dyscolus dejeanii
      • Dyscolus striatopunctatus
      • Dyscolus moreti
      • Dyscolus saxatilis
      • Dyscolus batallon
      • Dyscolus rotundatulus
      • Dyscolus opalescens
      • Dyscolus davidsoni
      • Dyscolus gratus
      • Dyscolus lagunensis
      • Dyscolus nitidulus
      • Dyscolus moritzi
      • Dyscolus incomis
      • Dyscolus degallieri
      • Dyscolus barbarellus
      • Dyscolus gemellus
      • Dyscolus robiginosus
      • Dyscolus ripicola
      • Dyscolus lherminieri
      • Dyscolus politus
      • Dyscolus alpinus
      • Dyscolus batesi
      • Dyscolus elongatus
      • Dyscolus dupuisi
      • Dyscolus lojaensis
      • Dyscolus immodicus
      • Dyscolus dominicensis
      • Dyscolus altarensis
      • Dyscolus desultor
      • Dyscolus segnipes
      • Dyscolus cristensis
      • Dyscolus cachetes
      • Dyscolus exsul
      • Dyscolus elliptolellus
      • Dyscolus alpaeoides
      • Dyscolus bliteus
      • Dyscolus carbonescens
      • Dyscolus margaritulus
      • Dyscolus obscurus
      • Dyscolus carbonarius
      • Dyscolus quadridentatus
      • Dyscolus involucer
      • Dyscolus redondensis
      • Dyscolus marginicollis
      • Dyscolus matucana
      • Dyscolus megacephalus
      • Dyscolus minusculus
      • Dyscolus bispinis
      • Dyscolus rufiventris
      • Dyscolus camur
      • Dyscolus ahenonotus
      • Dyscolus darwini
      • Dyscolus quadricollis
      • Dyscolus laevipennoides
      • Dyscolus jimenezi
      • Dyscolus fallax
      • Dyscolus rufulus
      • Dyscolus cyrtonotoides
      • Dyscolus umerangulatus
      • Dyscolus subcyaneus
      • Dyscolus mucuy
      • Dyscolus subviolaceus
      • Dyscolus onorei
      • Dyscolus laevilateris
      • Dyscolus bousqueti
      • Dyscolus spinipennis
      • Dyscolus lamottei
      • Dyscolus contumaza
      • Dyscolus cochabamba
      • Dyscolus brachyderus
      • Dyscolus gracilis
      • Dyscolus interruptus
      • Dyscolus protensus
      • Dyscolus brocchus
      • Dyscolus venezolanus
      • Dyscolus cordillerensis
      • Dyscolus albemarli
      • Dyscolus acutulus
      • Dyscolus quitensis
      • Dyscolus striatulus
      • Dyscolus aphaedrus
      • Dyscolus parviceps
      • Dyscolus yanacochae
      • Dyscolus villavicencio
      • Dyscolus arcanus
      • Dyscolus bogotensis
      • Dyscolus fusipalpis
      • Dyscolus buckleyi
      • Dyscolus antoninus
      • Dyscolus rectilineus
      • Dyscolus crabbei
      • Dyscolus festinus
      • Dyscolus leptomorphus
      • Dyscolus crassus
      • Dyscolus delenitor
      • Dyscolus viridiauratus
      • Dyscolus marginissimus
      • Dyscolus araneus
      • Dyscolus cozier
      • Dyscolus conicus
      • Dyscolus furvus
      • Dyscolus sinuosus
      • Dyscolus palatus
      • Dyscolus diopsis
      • Dyscolus ampliatus
      • Dyscolus orbicollis
      • Dyscolus patroboides
      • Dyscolus lucilius
      • Dyscolus nubilus
      • Dyscolus auyanensis
      • Dyscolus riveti
      • Dyscolus fronto
      • Dyscolus macerrimus
      • Dyscolus aemulus
      • Dyscolus bicolor
      • Dyscolus robustulus
      • Dyscolus mongusi
      • Dyscolus interstitialis
      • Dyscolus chalconotus
      • Dyscolus pallidipes
      • Dyscolus cyanellus
      • Dyscolus noctuabundus
      • Dyscolus pectoralis
      • Dyscolus otavaloensis
      • Dyscolus obesulus
      • Dyscolus paramemnonius
      • Dyscolus cyanicollis
      • Dyscolus princeps
      • Dyscolus erythrocerus
      • Dyscolus hemicyclicus
      • Dyscolus convergens
      • Dyscolus feronioides
      • Dyscolus hexacoelus
      • Dyscolus micans
      • Dyscolus nugax
      • Dyscolus thiemei
      • Dyscolus moestus
      • Dyscolus coloradito
      • Dyscolus moretianus
      • Dyscolus subauratus
      • Dyscolus asphaltinus
      • Dyscolus pollens
      • Dyscolus alternans
      • Dyscolus angustulus
      • Dyscolus palmae
      • Dyscolus macroderus
      • Dyscolus morosus
      • Dyscolus lyrophorus
      • Dyscolus nyctimus
      • Dyscolus sciakyi
      • Dyscolus breviculus
      • Dyscolus cruzensis
      • Dyscolus scabricollis
      • Dyscolus donrwi
      • Dyscolus cyanides
      • Dyscolus luciae
      • Dyscolus fratellus
      • Dyscolus bordoni
      • Dyscolus capsarius
      • Dyscolus callanga
      • Dyscolus imbaburae
      • Dyscolus portentosus
      • Dyscolus rotundiceps
      • Dyscolus omaseoides
      • Dyscolus trapezicollis
      • Dyscolus obuncus
      • Dyscolus subiridescens
      • Dyscolus caerulipennis
      • Dyscolus acuminatus
      • Dyscolus patocochae
      • Dyscolus oreas
      • Dyscolus cordicollis
      • Dyscolus memnonius
      • Dyscolus piceolus
      • Dyscolus deuvei
      • Dyscolus agonoides
      • Dyscolus falli
      • Dyscolus steropoides
      • Dyscolus eudemus
      • Dyscolus cordatus
      • Dyscolus ignicauda
      • Dyscolus angulosus
      • Dyscolus nevermanni
      • Dyscolus russeus
      • Dyscolus caucaensis
      • Dyscolus verecundus
      • Dyscolus superbus
      • Dyscolus glaucipennis
      • Dyscolus zunilensis
      • Dyscolus melanocnemis
      • Dyscolus bolivianus
      • Dyscolus prolixus
      • Dyscolus morenensis
      • Dyscolus raveloi
      • Dyscolus gaujoni
      • Dyscolus fartilis
      • Dyscolus porrectus
      • Dyscolus rutilans
      • Dyscolus transversicollis
      • Dyscolus mirandus
      • Dyscolus purpurellus
      • Dyscolus mucronatus
      • Dyscolus guatemalensis
      • Dyscolus ruficornis
      • Dyscolus semirufus
      • Dyscolus valens
      • Dyscolus beryllinus
      • Dyscolus lionotus
      • Dyscolus prostomis
      • Dyscolus viridans
      • Dyscolus nevadicus
      • Dyscolus platynellus
      • Dyscolus whymperi
      • Dyscolus haptoderoides
      • Dyscolus nocticolor
      • Dyscolus smithersi
      • Dyscolus velox
      • Dyscolus irriguus
      • Dyscolus ovatus
      • Dyscolus nitidus
      • Dyscolus biovatus
      • Dyscolus metallosomus
      • Dyscolus oopteroides
      • Dyscolus sulcatus
      • Dyscolus denigratus
      • Dyscolus delicatulus
      • Dyscolus purpuratus
      • Dyscolus giachinoi
      • Dyscolus mateui
      • Dyscolus acutus
      • Dyscolus despiciendus
      • Dyscolus iricolor
      • Dyscolus cuprascens
      • Dyscolus pseudoconicus
      • Dyscolus tachiranus
      • Dyscolus melanius
      • Dyscolus horni
      • Dyscolus lissomus
      • Dyscolus lubricus
      • Dyscolus brevipennis
      • Dyscolus inconspicuus
      • Dyscolus intergeneus
      • Dyscolus flavomarginatus
      • Dyscolus hirsutus
      • Dyscolus laetiusculus
      • Dyscolus olivaceus
      • Dyscolus acutipennis
      • Dyscolus cursor
      • Dyscolus pullatus
      • Dyscolus omissus
      • Dyscolus imitativus
      • Dyscolus tiguensis
      • Dyscolus durangensis
      • Dyscolus minimus
      • Dyscolus proteinus
      • Dyscolus virescens
      • Dyscolus dilutus
      • Dyscolus punctinotus
      • Dyscolus lamotteioides
      • Dyscolus bolivianoides
      • Dyscolus amethystinus
      • Dyscolus procephalus
      • Dyscolus unipunctatus
      • Dyscolus nebrianus
      • Dyscolus ovatulus
      • Dyscolus crypticulus
      • Dyscolus sellularius
      • Dyscolus dyschromus
      • Dyscolus orthomus
      • Dyscolus severus
      • Dyscolus reflexus
      • Dyscolus impiger
      • Dyscolus casalei
      • Dyscolus tenuicornis
      • Dyscolus montufari
      • Dyscolus monterredonda
      • Dyscolus woldai
      • Dyscolus fucatus
      • Dyscolus chaudoiri
      • Dyscolus oopterus
      • Dyscolus steinheili
      • Dyscolus limbicollis
      • Dyscolus montivagus
      • Dyscolus championi
      • Dyscolus stenophthalmus
      • Dyscolus caeruleomarginatus
      • Dyscolus unilobatus
      • Dyscolus cyanonotus
      • Dyscolus picicornis
      • Dyscolus lucidus
      • Dyscolus jelskii
      • Dyscolus subangulatus
      • Dyscolus agonellus
      • Dyscolus simoni
      • Dyscolus plebeius
      • Dyscolus columbinus
      • Dyscolus laticollis
      • Dyscolus atkinsi
      • Dyscolus troyaensis
      • Dyscolus lugens
      • Dyscolus joseensis
      • Dyscolus atratus
      • Dyscolus chrysopterus
      • Dyscolus funereus
      • Dyscolus melas
      • Dyscolus chontalensis
      • Dyscolus culatensis
      • Dyscolus ambagiosus
      • Dyscolus semiopacus
      • Dyscolus brachypterus
      • Dyscolus monachus
      • Dyscolus coptoderoides
      • Dyscolus grandicollis
      • Dyscolus meridanus
      • Dyscolus rubidus
      • Dyscolus castanipes
      • Dyscolus cleanthes
      • Dyscolus cyphopterus
      • Dyscolus segregatus
      • Dyscolus muzo
      • Dyscolus capticius
      • Dyscolus polylepidium
      • Dyscolus cyanostolus
      • Dyscolus phaeocnemis
      • Dyscolus atroaeneus
      • Dyscolus arvalis
      • Dyscolus evanescens
      • Dyscolus sublaevipennis
      • Dyscolus aequatorius
      • Dyscolus sellensis
      • Dyscolus viridipennis
      • Dyscolus subreflexus
      • Dyscolus algidus
      • Dyscolus sexfoveolatus
      • Dyscolus brunnellus
      • Dyscolus leleupi
      • Dyscolus cephalotes
      • Dyscolus lacertosus
      • Dyscolus chiriquinus
      • Dyscolus agilis
      • Dyscolus seriepunctatus
      • Dyscolus schunkei
      • Dyscolus variabilis
      • Dyscolus baorucensis
      • Dyscolus chathami
      • Dyscolus mucubajii
      • Dyscolus lutarius
      • Dyscolus ciliatus
      • Dyscolus ecuadoriensis
      • Dyscolus brullei
      • Dyscolus megalops
      • Dyscolus quadrilaterus
      • Dyscolus mucunuque
      • Dyscolus femoralis
      • Dyscolus hapax
      • Dyscolus hebeculus
      • Dyscolus bispinosus
      • Dyscolus cycloderus
      • Dyscolus laetificus
      • Dyscolus caeruleus
      • Dyscolus lignicola
      • Dyscolus platysmoides
      • Dyscolus caulatus
      • Dyscolus pristonychoides
      • Dyscolus furcillatus
      • Dyscolus macrous
      • Dyscolus lucifugus
      • Dyscolus platyderoides
      • Dyscolus epilissus
      • Dyscolus violaceipennis
      • Dyscolus capito
      • Dyscolus etontii
      • Dyscolus imitator
      • Dyscolus frigidus
      • Dyscolus teter
      • Dyscolus reichei
      • Dyscolus landolti
      • Dyscolus pyrophilus
      • Dyscolus spinifer
      • Dyscolus ebeninus
      • Dyscolus machetellus
      • Dyscolus reflexicollis
      • Dyscolus punoensis
      • Dyscolus concisus
      • Dyscolus latidens
      • Dyscolus maleodoratus
      • Dyscolus validus
      • Dyscolus drusillus
      • Dyscolus aeneipennis
      • Dyscolus physopterus
      • Dyscolus consanguineus
      • Dyscolus phaeolomus
      • Dyscolus deyrollei
      • Dyscolus incultus
      • Dyscolus austerus
      • Dyscolus vegaensis
      • Dyscolus tapiarius
      • Dyscolus ruminahui
      • Dyscolus tinctipennis
      • Dyscolus bucculentus
      • Dyscolus expansus
      • Dyscolus decorellus
      • Dyscolus defrictus
      • Dyscolus lamprotus
      • Dyscolus fragilis
      • Dyscolus rugulellus
      • Dyscolus albarragas
      • Dyscolus corvinus
      • Dyscolus trossulus
      • Dyscolus aterrimus
      • Dyscolus platynoides
      • Dyscolus tovarensis
      • Dyscolus cazieri
      • Dyscolus transfuga
    • Blackburnia → (141 species)
      • Blackburnia howarthi
      • Blackburnia ambiens
      • Blackburnia mandibularis
      • Blackburnia komohana
      • Blackburnia aaae
      • Blackburnia optimus
      • Blackburnia calathiformis
      • Blackburnia cheloniceps
      • Blackburnia paludicola
      • A Ground Beetle (Blackburnia cuneipennis)
      • Blackburnia kavanaughi
      • Blackburnia platyophthalmica
      • Blackburnia micans
      • Blackburnia abax
      • Blackburnia tibialis
      • Blackburnia haleakala
      • Blackburnia hihia
      • Blackburnia ewingi
      • Blackburnia hakeakapa
      • Blackburnia depressa
      • Blackburnia opaca
      • Blackburnia maculata
      • Blackburnia rupicola
      • Blackburnia rugosa
      • Blackburnia moerens
      • Blackburnia pilikua
      • Blackburnia concolor
      • Blackburnia atra
      • Blackburnia medeirosi
      • Blackburnia munroi
      • Blackburnia menehune
      • Blackburnia molokaiensis
      • Blackburnia muscicola
      • A Ground Beetle (Blackburnia kamehameha)
      • Blackburnia longula
      • Blackburnia godzilla
      • Blackburnia kilauea
      • Blackburnia cephalotes
      • Blackburnia bryophila
      • Blackburnia metromenoides
      • Blackburnia fulgida
      • Blackburnia asquithi
      • Blackburnia filipes
      • Blackburnia audax
      • Blackburnia kauwa
      • Blackburnia caliginosa
      • Blackburnia pauma
      • A Ground Beetle (Blackburnia lenta)
      • Blackburnia epicura
      • Blackburnia vagans
      • Blackburnia tantalus
      • Blackburnia calathoides
      • Blackburnia erythropus
      • Blackburnia polipoli
      • Blackburnia bartletti
      • Blackburnia perkinsi
      • Blackburnia huhula
      • Blackburnia mothra
      • Blackburnia sulcipennis
      • Blackburnia ovata
      • Blackburnia burneyi
      • Blackburnia cryptipes
      • Blackburnia ulaula
      • Blackburnia puncticeps
      • Blackburnia curtipes
      • A Ground Beetle (Blackburnia derodera)
      • Blackburnia corrusca
      • Blackburnia foveolata
      • Blackburnia gracilis
      • Blackburnia proterva
      • Blackburnia fracta
      • Blackburnia microps
      • Blackburnia polhemusi
      • Blackburnia erro
      • Blackburnia debilis
      • Blackburnia ipu
      • Blackburnia terebrata
      • Blackburnia fugitiva
      • Blackburnia costata
      • Blackburnia longipes
      • Blackburnia constricta
      • Blackburnia hilaris
      • Blackburnia lanaihalensis
      • A Ground Beetle (Blackburnia oceanica)
      • Blackburnia latifrons
      • Blackburnia aterrima
      • Blackburnia sphodriformis
      • Blackburnia fractistriata
      • Blackburnia viridis
      • Blackburnia fraterna
      • Blackburnia posticata
      • Blackburnia meticulosa
      • Blackburnia fraudator
      • Blackburnia agonoides
      • Blackburnia pavida
      • Blackburnia dyscolea
      • Blackburnia mystica
      • Blackburnia mutabilis
      • Blackburnia blaptoides
      • Blackburnia incendiaria
      • Blackburnia fordi
      • Blackburnia octoocellata
      • Blackburnia lata
      • Blackburnia punticeps
      • Blackburnia fossipennis
      • Blackburnia riparia
      • Blackburnia lanaiensis
      • Blackburnia kukui
      • Blackburnia elegans
      • Oahu Blackburnia ground beetle (Blackburnia insignis)
      • Blackburnia kipahulu
      • A Ground Beetle (Blackburnia transiens)
      • Blackburnia brevipes
      • Blackburnia kahili
      • Blackburnia lihau
      • Blackburnia palmae
      • Blackburnia frigida
      • Blackburnia waialeale
      • Blackburnia hawaiiensis
      • Blackburnia lucipetens
      • Blackburnia putealis
      • Blackburnia anomala
      • Blackburnia agilis
      • Blackburnia auana
      • Blackburnia abaxoides
      • Blackburnia sharpi
      • A Ground Beetle (Blackburnia barda)
      • Blackburnia pukalaina
      • Blackburnia gastrellariformis
      • Blackburnia perpolita
      • Blackburnia paloloensis
      • Blackburnia insociabilis
      • Blackburnia optata
      • Blackburnia limbata
      • Blackburnia kuiki
      • Blackburnia kauaiensis
      • Blackburnia koebelei
      • Blackburnia platynoides
      • Blackburnia tricolor
      • Blackburnia alternans
      • A Ground Beetle (Blackburnia micantipennis)
    • Tanystoma → (5 species)
      • Tanystoma cuyama
      • Tanystoma diabolicum
      • Tanystoma sulcatum
      • Tanystoma maculicolle
      • Tanystoma striatum
    • Metacolpodes → (31 species)
      • Metacolpodes incertus
      • Metacolpodes fryi
      • Metacolpodes godavaricus
      • Metacolpodes olivius
      • Metacolpodes amoenulus
      • Metacolpodes janakpurensis
      • Metacolpodes grandis
      • Metacolpodes parallelus
      • Metacolpodes janellus
      • Metacolpodes landrungensis
      • Metacolpodes janelloides
      • Metacolpodes hopkinsi
      • Metacolpodes cyaneus
      • Metacolpodes monticola
      • Metacolpodes buchanani
      • Metacolpodes rambouseki
      • Metacolpodes rotundatus
      • Metacolpodes tetraglochis
      • Metacolpodes superlita
      • Metacolpodes planithorax
      • Metacolpodes limodromoides
      • Metacolpodes laticeps
      • Metacolpodes rotundicollis
      • Metacolpodes truncatellus
      • Metacolpodes buxtoni
      • Metacolpodes buchannani
      • Metacolpodes laetus
      • Metacolpodes deliasianum
      • Metacolpodes hardwickii
      • Metacolpodes nilgherriensis
      • Metacolpodes takakuwai
    • Sericoda → (9 species)
      • A Ground Beetle (Sericoda quadripunctata)
      • Sericoda montana
      • A Ground Beetle (Sericoda bembidioides)
      • A Ground Beetle (Sericoda obsoleta)
      • Sericoda variolata
      • Sericoda ceylonica
      • Sericoda lissoptera
      • Sericoda balli
      • A Ground Beetle (Sericoda bogemannii)
    • Olisthopus → (21 species)
      • A Ground Beetle (Olisthopus micans)
      • Olisthopus brevicornis
      • Olisthopus glabricollis
      • Ground beetle (Olisthopus parmatus)
      • Olisthopus rotundatus
      • Olisthopus filicornis
      • Olisthopus pusio
      • Olisthopus iterans
      • Olisthopus glabratus
      • Olisthopus elongatus
      • Olisthopus humerosus
      • Olisthopus fuscatus
      • Olisthopus elburzensis
      • Olisthopus sturmii
      • Olisthopus inclavatus
      • Olisthopus palmensis
      • Olisthopus sibiricus
      • Olisthopus hispanicus
      • A Ground Beetle (Olisthopus innuens)
      • Olisthopus maderensis
      • Olisthopus ericae
    • Atranus → (2 species)
      • A Ground Beetle (Atranus pubescens)
      • Atranus ruficollis
    • Rhadine → (56 species)
      • A Cave Obligate Beetle (Rhadine austinica)
      • Rhadine nr. chipinque WMoore DNA3088
      • Rhadine euprepes
      • Rhadine umbra
      • Rhadine bullis
      • Rhadine longiceps
      • Rhadine sprousei
      • Rhadine specum
      • A Ground Beetle (Rhadine tenebrosa)
      • Rhadine bolivari
      • Rhadine medellini
      • Rhadine leptodes
      • A Ground Beetle (Rhadine caudata)
      • Rhadine jejuna
      • Rhadine nivalis
      • Rhadine persephone
      • Rhadine pelaezi
      • A Cave Obligate Beetle (Rhadine russelli)
      • Rhadine nr. austinica WMoore DNA3061
      • Rhadine testacea
      • Rhadine perlevis
      • Rhadine pertenuis
      • A Ground Beetle (Rhadine infernalis)
      • Rhadine exilis
      • Rhadine rossi
      • A Ground Beetle (Rhadine koepkei)
      • Rhadine longicollis
      • Rhadine chipinque
      • Rhadine dissecta
      • Rhadine ivyi
      • Rhadine nr. longicollis WMoore DNA3044
      • Rhadine rubra
      • Rhadine howdeni
      • Rhadine albamontana
      • Rhadine elliotti
      • Rhadine testacaea
      • Rhadine constricta
      • Rhadine myrmecodes
      • Rhadine insolita
      • Rhadine balesi
      • Rhadine sublustris
      • Rhadine araizai
      • A Ground Beetle (Rhadine subterranea)
      • Rhadine lanei
      • Rhadine reyesi
      • Rhadine pugetana
      • Rhadine longipes
      • Rhadine reddelli
      • A Cave Obligate Beetle (Rhadine noctivaga)
      • A Ground Beetle (Rhadine lindrothi)
      • Rhadine anthicoides
      • A Ground Beetle (Rhadine ozarkensis)
      • Rhadine hendrichsi
      • Rhadine rotgeri
      • Rhadine larvalis
      • Rhadine grubbsi
    • Incagonum → (24 species)
      • Incagonum andicola
      • Incagonum lineatopunctatum
      • Incagonum ambiguum
      • Incagonum discosulcatum
      • Incagonum lucidum
      • Incagonum pedestre
      • Incagonum peruvianum
      • Incagonum cordicolle
      • Incagonum semistriatum
      • Incagonum opacum
      • Incagonum gayi
      • Incagonum chilense
      • Incagonum circumdatum
      • Incagonum inca
      • Incagonum dejeani
      • Incagonum bonariense
      • Incagonum mateui
      • Incagonum laevicolle
      • Incagonum angulatum
      • Incagonum hiekei
      • Incagonum brasiliense
      • Incagonum melas
      • Incagonum quadricolle
      • Incagonum aeneum
    • Anchomenus → (20 species)
      • Anchomenus aeneolus
      • Anchomenus pictonensis
      • Anchomenus funebris
      • Anchomenus dohrnii
      • Anchomenus capensis
      • Anchomenus virescens
      • Anchomenus turkestanicus
      • Anchomenus punctulatus
      • Anchomenus bipunctatus
      • Anchomenus leucopus
      • Anchomenus bellus
      • Anchomenus alcedo
      • Anchomenus suensoni
      • Anchomenus kurnakovi
      • Anchomenus cyaneus
      • Anchomenus dorsalis
      • A Ground Beetle (Anchomenus quadratus)
      • Anchomenus yukihikoi
      • Anchomenus orphanus
      • Anchomenus kataevi
    • Liagonum → (55 species)
      • Liagonum viettei
      • Liagonum ranomandryae
      • Liagonum deplanatum
      • Liagonum anosyanum
      • Liagonum baleense
      • Liagonum peyrierasi
      • Liagonum incertum
      • Liagonum sexpunctatum
      • Liagonum chappuisi
      • Liagonum vicinum
      • Liagonum nebrioides
      • Liagonum florens
      • Liagonum bicolor
      • Liagonum scordiscum
      • Liagonum pauliani
      • Liagonum kiymbiae
      • Liagonum decellei
      • Liagonum solidum
      • Liagonum vakoanae
      • Liagonum analavelonae
      • Liagonum hova
      • Liagonum fulvipes
      • Liagonum orophilum
      • Liagonum vadoni
      • Liagonum bosmansi
      • Liagonum aereum
      • Liagonum submimum
      • Liagonum marakwetianum
      • Liagonum monticola
      • Liagonum vanderijsti
      • Liagonum curvipes
      • Liagonum scotti
      • Liagonum metrium
      • Liagonum bintumanum
      • Liagonum marojejyanum
      • Liagonum mantasoae
      • Liagonum laticolle
      • Liagonum mangindranum
      • Liagonum ueleanum
      • Liagonum simplex
      • Liagonum brachypterum
      • Liagonum coquereli
      • Liagonum descarpentriesi
      • Liagonum assimile
      • Liagonum chenzemae
      • Liagonum lichenyanum
      • Liagonum tsaratananae
      • Liagonum arecarum
      • Liagonum grandidieri
      • Liagonum mahafalyanum
      • Liagonum jeanneli
      • Liagonum rhetoborum
      • Liagonum mandibulare
      • Liagonum beckeri
      • Liagonum subsolanum
    • Neomegalonychus crassipes
  • Sphodrini → (719 species)
    • Laemostenus → (212 species)
      • Laemostenus kashmirensis
      • Laemostenus quadricollis
      • Laemostenus sciakyi
      • Laemostenus kurdicus
      • Laemostenus mateui
      • Laemostenus piceus
      • Laemostenus deneveui
      • Laemostenus jeannelianus
      • Laemostenus modestoides
      • Laemostenus mannerheimii
      • Laemostenus mogadoricus
      • Laemostenus kolbi
      • Laemostenus pisidicus
      • Laemostenus giachinoi
      • Laemostenus elburzensis
      • Laemostenus picicornis
      • Laemostenus fezzensis
      • Laemostenus peloponnesiacus
      • Laemostenus sulaimanensis
      • Laemostenus angustatus
      • Laemostenus kirschenhoferi
      • Laemostenus leonhardi
      • Laemostenus foucauldi
      • Laemostenus parallelocollis
      • Laemostenus zakariyiensis
      • Laemostenus leptoderus
      • Laemostenus vagabundus
      • Laemostenus praedictus
      • Laemostenus sericeus
      • Laemostenus esfandiarii
      • Laemostenus melillensis
      • Laemostenus obtusangulus
      • Laemostenus spinifer
      • Laemostenus monguzzii
      • Laemostenus morvani
      • Laemostenus koenigi
      • Laemostenus cazorlensis
      • Laemostenus conspicuus
      • Laemostenus petrimagni
      • Laemostenus gobbii
      • Laemostenus pakistanus
      • Laemostenus yazdensis
      • Laemostenus recticollis
      • Laemostenus bergvalli
      • Laemostenus lyauteyi
      • Laemostenus rudichae
      • Laemostenus euxinicus
      • Laemostenus zagrosensis
      • Laemostenus mairei
      • Laemostenus caussolensis
      • Laemostenus ponticola
      • Laemostenus venustus
      • Laemostenus stussineri
      • Laemostenus tiouirii
      • Laemostenus thessalicus
      • Laemostenus malhommei
      • Laemostenus sonkulensis
      • Laemostenus prolixus
      • Laemostenus macropus
      • Laemostenus martensianus
      • Laemostenus speleophilus
      • Laemostenus bhadarwahensis
      • Laemostenus caspius
      • Laemostenus ganglbauerianus
      • Laemostenus dalmatinus
      • Laemostenus alejandroi
      • Laemostenus gomezi
      • Laemostenus tacitus
      • Laemostenus elegans
      • Laemostenus moldoensis
      • Laemostenus villardi
      • Laemostenus nepalensis
      • Laemostenus demoflysi
      • Laemostenus troglophilus
      • Laemostenus casalei
      • Laemostenus berberus
      • Laemostenus peleus
      • Laemostenus janczyki
      • Laemostenus iranicus
      • Laemostenus levis
      • Laemostenus subtilissimus
      • Laemostenus turkestanicus
      • Laemostenus colossus
      • Laemostenus gasparoi
      • Laemostenus tenuis
      • Laemostenus andreevi
      • Laemostenus mauritanicus
      • Laemostenus asiaeminoris
      • Laemostenus amasiae
      • Laemostenus favieri
      • Laemostenus glasunowi
      • Laemostenus atlanticus
      • Laemostenus alluaudi
      • Laemostenus mirzayani
      • Laemostenus cadilhaci
      • Laemostenus barrancoi
      • Laemostenus oblongus
      • Laemostenus longicornis
      • Laemostenus janthinus
      • Laemostenus migliaccioi
      • Laemostenus brunneus
      • Laemostenus lestes
      • Laemostenus reissi
      • Laemostenus andalusiacus
      • Laemostenus balkenohli
      • Laemostenus krueperi
      • Laemostenus algerinus
      • Laemostenus latialis
      • Laemostenus bulirschi
      • Laemostenus lundbergi
      • Laemostenus schrammi
      • Laemostenus jailensis
      • Laemostenus obtusus
      • Laemostenus zoiai
      • Laemostenus badakshanus
      • Laemostenus aelleni
      • Laemostenus bodemeyeri
      • cosmopolitan ground beetle (Laemostenus complanatus)
      • Laemostenus magellensis
      • Laemostenus chitralicus
      • Laemostenus levantinus
      • Laemostenus cavazzutii
      • Laemostenus quadrangulus
      • Laemostenus visai
      • Laemostenus insubricus
      • Laemostenus vignai
      • Laemostenus levushkini
      • Laemostenus brancuccii
      • Laemostenus schreibersii
      • Laemostenus aequalis
      • Laemostenus deconincki
      • Laemostenus derventicus
      • Laemostenus pokornyi
      • Laemostenus anomostriatus
      • Laemostenus alpinus
      • Laemostenus luristanus
      • Laemostenus catlosi
      • Laemostenus capito
      • Laemostenus dolini
      • Laemostenus cavicola
      • Laemostenus seguranus
      • Laemostenus nusayriyahensis
      • Laemostenus bermejae
      • Laemostenus suramensis
      • Laemostenus conradti
      • Laemostenus cyrenaicus
      • Laemostenus elytrolargus
      • Laemostenus ginellae
      • Laemostenus agnolettii
      • Laemostenus straneoi
      • Laemostenus heinzi
      • Laemostenus elongatus
      • Laemostenus pretiosus
      • Laemostenus actenipoides
      • Laemostenus nigrescens
      • Laemostenus ajmasianus
      • Laemostenus susicus
      • Laemostenus cordicollis
      • Laemostenus ferghanicus
      • Laemostenus cyrili
      • Laemostenus rigrevanensis
      • Laemostenus punctatostriatus
      • Laemostenus largomarginalis
      • Laemostenus guzelolukensis
      • Laemostenus dubaulti
      • Laemostenus pinicola
      • Laemostenus sanguinipes
      • Laemostenus libanensis
      • Laemostenus shirazensis
      • Laemostenus kabylicus
      • Laemostenus carinatus
      • Laemostenus kuraminensis
      • Laemostenus baehri
      • Laemostenus lederi
      • Laemostenus baeticus
      • Laemostenus tschitscherini
      • Laemostenus acutangulus
      • Laemostenus vidali
      • Laemostenus ledereri
      • Laemostenus wittmeri
      • Laemostenus hroni
      • Laemostenus gratus
      • Laemostenus tichyi
      • Laemostenus ponticus
      • Laemostenus rimmae
      • Laemostenus antonrichteri
      • Laemostenus beroni
      • Laemostenus stoevi
      • Laemostenus meaillensis
      • Laemostenus pseudobrunneus
      • Laemostenus extensus
      • Laemostenus puchneri
      • Laemostenus kestelensis
      • Laemostenus cimmerius
      • Laemostenus barbarus
      • Laemostenus ambroggii
      • Laemostenus lassallei
      • Laemostenus patrizii
      • Laemostenus arthuri
      • Laemostenus portsensis
      • Laemostenus pippiai
      • Laemostenus aegyptiacus
      • A Ground Beetle (Laemostenus terricola)
      • Laemostenus tentiobtusus
      • Laemostenus gwendyddae
      • Laemostenus navaricus
      • Laemostenus capitatus
      • Laemostenus loudai
      • Laemostenus anatolicus
      • Laemostenus plasoni
      • Laemostenus farsicus
      • Laemostenus bicolor
    • Calathus → (179 species)
      • Lauricalathus → (19 species)
        • Calathus angustulus
        • Calathus cognatus
        • Calathus appendiculatus
        • Calathus vividus
        • Calathus gomerensis
        • Calathus abaxoides
        • Calathus auctus
        • Calathus freyi
        • Calathus canariensis
        • Calathus spretus
        • Calathus depressus
        • Calathus carinatus
        • Calathus ascendens
        • Calathus colasianus
        • Calathus rectus
        • Calathus laurenticola
        • Calathus angularis
        • Calathus marcellae
        • Calathus complanatus
      • Calathus subgenus Calathus → (28 species)
        • Calathus moralesi
        • Calathus fracassii
        • Calathus malacensis
        • A Ground Beetle (Calathus fuscipes)
        • Calathus arcuatus
        • Calathus oreades
        • Calathus zabroides
        • Calathus syriacus
        • Calathus busii
        • Calathus mirei
        • Calathus distinguendus
        • Calathus hispanicus
        • Calathus montivagus
        • Calathus korax
        • Calathus roccai
        • Calathus minutus
        • Calathus vivesi
        • Calathus ellipticus
        • Calathus uniseriatus
        • Calathus glabricollis
        • Calathus erzeliki
        • Calathus baeticus
        • Calathus longicollis
        • Calathus giganteus
        • Calathus luctuosus
        • Calathus sirentensis
        • Calathus libanensis
        • Calathus vuillefroyi
      • Trichocalathus → (3 species)
        • Calathus pilosipennis
        • Calathus obliteratus
        • Calathus refleximargo
      • Amphyginus → (4 species)
        • Calathus semisericeus
        • Calathus piceus
        • Calathus rotundicollis
        • Calathus rotundatus
      • Neocalathus → (14 species)
        • Calathus granatensis
        • Calathus deplanatus
        • Calathus micropterus
        • Calathus gonzalezi
        • A Ground Beetle (Calathus ruficollis)
        • Calathus ambiguus
        • Calathus mollis
        • Calathus erratus
        • Calathus melanocephalus
        • Calathus metallicus
        • Calathus subfuscus
        • Calathus cinctus
        • Calathus simplicicollis
        • Calathus asturiensis
      • Calathus tombesii
      • Calathus scotti
      • Calathus kryzhanovskii
      • Calathus gelascens
      • Calathus gansuensis
      • Calathus vagestriatus
      • Calathus theodori
      • Calathus ras
      • Calathus horsti
      • Calathus bosnicus
      • Calathus annapurnae
      • Calathus balli
      • Calathus clauseni
      • Calathus parvicollis
      • Calathus oertzeni
      • Calathus albanicus
      • Calathus baleensis
      • Calathus rufocastaneus
      • Calathus lissoderus
      • Calathus rhaticus
      • Calathus aethiops
      • Calathus kirschenhoferianus
      • A Ground Beetle (Calathus calceus)
      • Calathus ambigens
      • Calathus relictus
      • Calathus jakupicensis
      • Calathus suffuscus
      • Calathus durango
      • Calathus aeneocupreus
      • Calathus erwini
      • Calathus carvalhoi
      • Calathus shoanus
      • Calathus nitidus
      • Calathus acuticollis
      • Calathus chokeensis
      • Calathus guadalupensis
      • Calathus kollari
      • Calathus robustus
      • Calathus reflexus
      • Calathus sotshiensis
      • Calathus himalayae
      • Calathus chioriae
      • Calathus montanellus
      • Calathus vicenteorum
      • Calathus jacupicensis
      • Calathus oreobius
      • Calathus hyrcanus
      • Calathus kirschenhoferi
      • Calathus scottianus
      • Calathus wolloensis
      • Calathus vignatagliantii
      • Calathus orthomoides
      • Calathus rubripes
      • Calathus laticaudis
      • Calathus grandiceps
      • Calathus manasluensis
      • Calathus heinertzi
      • Calathus praestans
      • Calathus grandicollis
      • Calathus alternans
      • Calathus ollivieri
      • Calathus montanus
      • Calathus holzschuhi
      • Calathus peltatus
      • Calathus atitari
      • Calathus trapezicollis
      • Calathus femoralis
      • Calathus ignicollis
      • Calathus kebedei
      • Calathus leptodactylus
      • Calathus mandibularis
      • Calathus ankoberensis
      • Calathus fimbriatus
      • Calathus pecoudi
      • Calathus stricticaudis
      • Calathus extensicollis
      • Calathus wittmerianus
      • Calathus patrizii
      • Calathus deyrollei
      • Calathus ravasinii
      • Calathus algens
      • Calathus reflexicollis
      • Calathus carballalae
      • Calathus reebae
      • Calathus heinzianus
      • Calathus afroalpinus
      • Calathus whiteheadi
      • Calathus gunaensis
      • Calathus queinneci
      • Calathus caucasicus
      • Calathus subpraestans
      • Calathus macedonicus
      • Calathus casalei
      • Calathus anatolicus
      • Calathus gugheensis
      • Calathus deliae
      • Calathus elpis
      • Calathus anistschenkoi
      • Calathus juan
      • Calathus focarilei
      • Calathus lundbladi
      • Calathus simienensis
      • Calathus pommeranzi
      • Calathus ovipennis
      • Calathus recticaudis
      • Calathus dubaulti
      • Calathus rotgeri
      • Calathus potosi
      • Calathus circumseptus
      • Calathus amplius
      • Calathus ciliatus
    • Dolichus → (3 species)
      • Dolichus davidis
      • Dolichus halensis
      • Dolichus coreicus
    • Synuchus → (82 species)
      • Synuchus ventricosus
      • Synuchus satoi
      • Synuchus bellus
      • Synuchus nordmanni
      • Synuchus cycloderus
      • Synuchus keinigus
      • Synuchus hummeli
      • Synuchus calathinus
      • Synuchus masumotoi
      • Synuchus brevis
      • Synuchus adelosia
      • Synuchus robustus
      • Synuchus fukuharai
      • Synuchus microtes
      • Synuchus chabo
      • Synuchus picicolor
      • Synuchus himalayicus
      • Synuchus intermedius
      • Synuchus semirufus
      • Synuchus hikosanus
      • Synuchus arcuaticollis
      • Synuchus sinomeridionalis
      • Synuchus laticollis
      • Synuchus sichuanensis
      • Synuchus inadai
      • Synuchus callitheres
      • Synuchus gigas
      • Synuchus montanus
      • Synuchus limbalis
      • Synuchus macer
      • Synuchus narae
      • Synuchus rjabuchini
      • Synuchus dulcigradus
      • Synuchus formosanus
      • Synuchus rufulus
      • Synuchus ishigakiensis
      • Synuchus pinguiusculus
      • Synuchus pseudomorphus
      • Synuchus minimus
      • Synuchus agonus
      • Synuchus angustus
      • Synuchus taiwanus
      • Synuchus fulvus
      • Synuchus truncatus
      • Synuchus rufofuscus
      • Synuchus longissimus
      • Synuchus suensoni
      • Synuchus sinuaticollis
      • Synuchus gravidus
      • Synuchus pulcher
      • Synuchus crocatus
      • Synuchus nitidus
      • Synuchus tanzawanus
      • Synuchus vivalis
      • Synuchus coreanus
      • Synuchus nanpingensis
      • Synuchus sikkimensis
      • Synuchus major
      • Synuchus rectangulus
      • Synuchus melantho
      • Synuchus patroboides
      • Synuchus breviusculus
      • Synuchus amamioshimae
      • Synuchus elburzensis
      • Synuchus shibatai
      • Synuchus pallidulus
      • Synuchus tokararum
      • Synuchus pallipes
      • Synuchus agonoides
      • Ground beetle (Synuchus impunctatus)
      • Synuchus angusticeps
      • Synuchus longipes
      • Synuchus chinensis
      • Synuchus atricolor
      • Synuchus testaceus
      • Synuchus congruus
      • Synuchus andrewesi
      • Synuchus dubius
      • Synuchus assamensis
      • Synuchus takeuchii
      • Synuchus cathaicus
      • Synuchus yasumatsui
    • Acalathus → (14 species)
      • Acalathus wrzecionkoi
      • Acalathus luhuoensis
      • Acalathus nanpingensis
      • Acalathus kangdingensis
      • Acalathus validulus
      • Acalathus langmusiensis
      • Acalathus businskyi
      • Acalathus shaanxiensis
      • Acalathus fallax
      • Acalathus semirufescens
      • Acalathus yunnanicus
      • Acalathus drolmae
      • Ground beetle (Acalathus advena)
      • Acalathus xiahensis
    • Lindrothius → (2 species)
      • Lindrothius caucasicus
      • Lindrothius pseudopraestans
    • Xestopus → (7 species)
      • Xestopus alticola
      • Xestopus kumatai
      • Xestopus nepalensis
      • Xestopus cyaneus
      • Xestopus bhutanensis
      • Xestopus walteri
      • Xestopus cordicollis
    • Platyderus → (111 species)
      • Platyderus alticola
      • Platyderus sagrensis
      • Platyderus beseanus
      • Platyderus speleus
      • Platyderus preciosae
      • Platyderus talyschensis
      • Platyderus leonensis
      • Platyderus gallaecus
      • Platyderus ortunoi
      • Platyderus graecus
      • Platyderus tadzhikistanus
      • Platyderus filicornis
      • Platyderus grandiceps
      • Platyderus calathoides
      • Platyderus sinensis
      • Platyderus juncoi
      • Platyderus quadricollis
      • Platyderus insignitus
      • Platyderus solissimus
      • Platyderus dalmatinus
      • Platyderus balearicus
      • Platyderus taghizadehi
      • Platyderus cyrtensis
      • Platyderus portalegrae
      • Platyderus vuillefroyi
      • Platyderus robustus
      • Platyderus torressalai
      • Platyderus moncayensis
      • Platyderus dejeani
      • Platyderus troglodytes
      • Platyderus aragonicus
      • Platyderus berlovorum
      • Platyderus otini
      • Platyderus davatchii
      • Platyderus corcyreus
      • Platyderus haberhaueri
      • Platyderus punctiger
      • Platyderus atticus
      • Platyderus cyprius
      • Platyderus canaliculatus
      • Platyderus jedlickai
      • Platyderus formenterae
      • Platyderus valencianus
      • Platyderus magrinii
      • Platyderus barsevskisi
      • Platyderus majoricus
      • Platyderus reticulatus
      • Platyderus itziarae
      • Platyderus asturiensis
      • Platyderus lusitanicus
      • Platyderus zagrosensis
      • Platyderus ledouxi
      • Platyderus anandi
      • Platyderus jeannei
      • Platyderus subcrenatus
      • Platyderus crypticola
      • Platyderus robustoides
      • Platyderus mateui
      • Platyderus ferrantei
      • Platyderus vignai
      • Platyderus testaceus
      • Platyderus varians
      • Platyderus gregarius
      • Platyderus gazureki
      • Platyderus migelangeli
      • Platyderus breuili
      • Platyderus montanellus
      • Platyderus barrosi
      • Platyderus languidus
      • Platyderus namrun
      • Platyderus toribioi
      • Platyderus dertosensis
      • Platyderus marduk
      • Platyderus elegans
      • Platyderus schuberti
      • Platyderus foveipennis
      • Platyderus rotundatus
      • Platyderus saezi
      • Platyderus anatolicus
      • Platyderus marianicus
      • Platyderus ellipticus
      • Platyderus espanoli
      • Platyderus emblema
      • Platyderus minutus
      • Platyderus neapolitanus
      • Platyderus sciakyi
      • Platyderus ruficollis
      • Platyderus vivesi
      • Platyderus umbratus
      • Platyderus ruizi
      • Platyderus zaballosi
      • Platyderus paganettii
      • Platyderus caucasicus
      • Platyderus casalei
      • Platyderus rufus
      • Platyderus notatus
      • Platyderus bolivari
      • Platyderus salmantinus
      • Platyderus lencinai
      • Platyderus algesiranus
      • Platyderus incertans
      • Platyderus weiratheri
      • Platyderus coiffaiti
      • Platyderus akkus
      • Platyderus quezeli
      • Platyderus lancerottensis
      • Platyderus skoupyi
      • Platyderus alhamillensis
      • Platyderus lombardii
      • Platyderus pyrenaeus
      • Platyderus chodjaii
    • Licinopsis → (6 species)
      • Licinopsis schurmanni
      • Licinopsis angustula
      • Licinopsis obliterata
      • Licinopsis picescens
      • Licinopsis gaudini
      • Licinopsis alternans
    • Anchomenidius → (2 species)
      • Anchomenidius astur
      • Anchomenidius feldmanni
    • Pristosia → (97 species)
      • Pristosia clara
      • Pristosia aquilo
      • Pristosia strigipennis
      • Pristosia vigil
      • Pristosia tenuistriata
      • Pristosia oblonga
      • Pristosia abaensis
      • Pristosia purpurea
      • Pristosia dongziensis
      • Pristosia meridionalis
      • Pristosia miwai
      • Pristosia xanthopus
      • Pristosia robusta
      • Pristosia lateritioides
      • Pristosia leurops
      • Pristosia leptodes
      • Pristosia crenata
      • Pristosia lacerans
      • Pristosia illustris
      • Pristosia nitouensis
      • Pristosia wenxianensis
      • Pristosia glacialis
      • Pristosia elevata
      • Pristosia bastai
      • Pristosia maoniushanensis
      • Pristosia morvani
      • Pristosia taibaishanensis
      • Pristosia magna
      • Pristosia impunctata
      • Pristosia suensoni
      • Pristosia reitteri
      • Pristosia heinzi
      • Pristosia chlorodes
      • Pristosia dahud
      • Pristosia acraea
      • Pristosia szekessyi
      • Pristosia wrasei
      • Pristosia cupreata
      • Pristosia jureceki
      • Pristosia chambae
      • Pristosia szetschuana
      • Pristosia heyrovskyi
      • Pristosia nitidula
      • Pristosia championi
      • Pristosia alesi
      • Pristosia hweisiensis
      • Pristosia aeneola
      • Pristosia latistoma
      • Pristosia proxima
      • Pristosia amaroides
      • Pristosia sulcipennis
      • Pristosia ledouxi
      • Pristosia falsicolor
      • Pristosia bulirschi
      • Pristosia qiaojiacensis
      • Pristosia pingwuensis
      • Pristosia yunnana
      • Pristosia similata
      • Pristosia quadricolor
      • Pristosia deqenensis
      • Pristosia maoxianensis
      • Pristosia schnelli
      • Pristosia jedlickai
      • Pristosia glabella
      • Pristosia braccata
      • Pristosia nubilipenissoides
      • Pristosia minutalis
      • Pristosia colasi
      • Pristosia flava
      • Pristosia sciakyi
      • Pristosia meiliensis
      • Pristosia nanpingicus
      • Pristosia striata
      • Pristosia atrema
      • Pristosia nubilipennis
      • Pristosia purkynei
      • Pristosia litangensis
      • Pristosia przewalskii
      • Pristosia sienla
      • Pristosia lateritia
      • Pristosia delavayi
      • Pristosia brancuccii
      • Pristosia hauseri
      • Pristosia zheduoensis
      • Pristosia picescens
      • Pristosia dodensis
      • Pristosia nepalensis
      • Pristosia silvanoi
      • Pristosia thilliezi
      • Pristosia wrzecionkoi
      • Pristosia acalathusoides
      • Pristosia aereipennis
      • Pristosia potanini
      • Pristosia tibetana
      • Pristosia viridis
      • Pristosia opaca
      • Pristosia sterbai
    • Amaroschema gaudini
    • Miquihuana rhadiniformis
    • Thermoscelis insignis
    • Synuchidius ganglbaueri

Other links

• Wikidata
• Wikipedia (language: en)
• Wikipedia (language: fr)
• Wikipedia (language: ru)
• Wikipedia (language: nl)
• Encyclopedia of Life
• NCBI (genetics)"
• Open Tree of Life

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