Base of the evolutionary tree for Pterostichus

← parent

• Pterostichus (1243 species)
  • Platypterus → (3 species)
    • Pterostichus panzeri
    • Pterostichus ziegleri
    • Pterostichus truncatus
  • Pseudomaseus → (11 species)
    • Pterostichus minor
    • Pterostichus rotundangulus
    • Pterostichus gracilis
    • Pterostichus ambigenus
    • Pterostichus rhaeticus
    • Pterostichus fuscicornis
    • Ground beetle (Pterostichus luctuosus)
    • Pterostichus anthracinus
    • Pterostichus chujoiellus
    • Pterostichus nigrita
    • Pterostichus tenuis
  • Ethira → (3 species)
    • Pterostichus multiseta
    • Pterostichus cometes
    • Pterostichus pilifer
  • Oreophilus → (6 species)
    • Pterostichus spinolae
    • Pterostichus morio
    • Pterostichus flavofemoratus
    • Pterostichus multipunctatus
    • Pterostichus bicolor
    • A Ground Beetle (Pterostichus jurinei)
  • Pterostichus subgenus Pterostichus → (6 species)
    • Pterostichus kyushuensis
    • Pterostichus hagenbachii
    • Pterostichus shiibanus
    • Pterostichus rutilans
    • Pterostichus cristatus
    • Pterostichus masidai
  • Badistrinus → (2 species)
    • Pterostichus bandotaro
    • Pterostichus polygenus
  • Morphnosoma → (4 species)
    • Ground beetle (Pterostichus melanarius)
    • Pterostichus robustistylis
    • Pterostichus habui
    • Pterostichus thunbergi
  • Paralianoe → (4 species)
    • Pterostichus bisetosus
    • Pterostichus daisenicus
    • Pterostichus biexcisus
    • Pterostichus uenoi
  • Epinialoe → (3 species)
    • Pterostichus dandonis
    • Pterostichus cristatoides
    • Pterostichus spiculifer
  • Eurythoracana → (2 species)
    • Pterostichus haptoderoides
    • Pterostichus kajimurai
  • Nialoe → (14 species)
    • Pterostichus abaciformis
    • Pterostichus shibatai
    • Pterostichus yakushimanus
    • Pterostichus brunneipennis
    • Pterostichus scurra
    • Pterostichus raptor
    • Pterostichus taoi
    • Pterostichus opacipennis
    • Pterostichus sincerus
    • Pterostichus praedo
    • Pterostichus seunglaki
    • Pterostichus plesiomorphus
    • Pterostichus asymmetricus
    • Pterostichus katashinensis
  • Phonias → (4 species)
    • Pterostichus diligens
    • A Ground Beetle (Pterostichus strenuus)
    • Pterostichus ovoideus
    • Pterostichus longinquus
  • Metallophilus → (2 species)
    • Pterostichus interruptus
    • A Ground Beetle (Pterostichus sublaevis)
  • Hypherpes subgenus complex → (61 species)
    • Hypherpes → (34 species)
      • Pterostichus diabolus
      • A Ground Beetle (Pterostichus algidus)
      • Pterostichus lattini
      • A Ground Beetle (Pterostichus herculaneus)
      • Pterostichus nr. ybousqueti KWW-2007
      • A Ground Beetle (Pterostichus castaneus)
      • Pterostichus serripes
      • Pterostichus congestus
      • Pterostichus nr. californicus KWW-1
      • Pterostichus nr. californicus KWW-2
      • Pterostichus isabellae
      • Pterostichus nr. crenicollis KWW-2007
      • A Ground Beetle (Pterostichus nigrocaeruleus)
      • Pterostichus brachylobus
      • A Ground Beetle (Pterostichus protractus)
      • Pterostichus intectus
      • Pterostichus nr. congestus KWW-1
      • Pterostichus miscellus
      • Pterostichus tuberculofemoratus
      • Pterostichus tarsalis
      • Pterostichus nr. congestus KWW-2
      • Pterostichus canallatus
      • A Ground Beetle (Pterostichus ecarinatus)
      • Pterostichus californicus
      • Pterostichus menetriesii
      • A Ground Beetle (Pterostichus tristis)
      • Pterostichus jacobinus
      • Pterostichus lama
      • A Ground Beetle (Pterostichus adoxus)
      • A Ground Beetle (Pterostichus amethystinus)
      • A Ground Beetle (Pterostichus neobrunneus)
      • Pterostichus inermis
      • Pterostichus morionides
      • A Ground Beetle (Pterostichus crenicollis)
    • Anilloferonia → (2 species)
      • Pterostichus lanei
      • Pterostichus testaceus
    • Leptoferonia → (25 species)
      • Pterostichus yosemitensis
      • Pterostichus sphodrinus
      • Pterostichus infernalis
      • Pterostichus inopinus
      • Pterostichus cochlearis
      • Pterostichus marinensis
      • A Ground Beetle (Pterostichus pumilus)
      • Pterostichus humilis
      • Pterostichus blodgettensis
      • Pterostichus beyeri
      • Pterostichus pemphredo
      • Pterostichus trinitensis
      • Pterostichus enyo
      • Pterostichus lobatus
      • Pterostichus fuchsi
      • Roth's blind ground beetle (Pterostichus rothi)
      • Pterostichus hatchi
      • Pterostichus idahoae
      • Pterostichus fenyesi
      • Pterostichus mattolensis
      • A Ground Beetle (Pterostichus inanis)
      • Pterostichus stapedius
      • Pterostichus caligans
      • Pterostichus angustus
      • Pterostichus hackerae
  • Sinosteropus → (2 species)
    • Pterostichus pseudojugivagus
    • Pterostichus pseudosinensis
  • Eosteropus → (6 species)
    • Pterostichus moestus
    • Pterostichus orientalis
    • Pterostichus japonicus
    • Pterostichus aethiops
    • Pterostichus karasawai
    • Pterostichus fuligineus
  • Lenapterus → (3 species)
    • A Ground Beetle (Pterostichus punctatissimus)
    • A Ground Beetle (Pterostichus agonus)
    • Pterostichus subrugosus
  • Lyrothorax → (3 species)
    • Pterostichus fujitai
    • Pterostichus yoritomus
    • Pterostichus caspius
  • Bothriopterus → (8 species)
    • Pterostichus abasarukini
    • Pterostichus quadrifoveolatus
    • Pterostichus subovatus
    • Ground beetle (Pterostichus adstrictus)
    • Pterostichus oblongopunctatus
    • Pterostichus aeneocupreus
    • Ground beetle (Pterostichus mutus)
    • Ground beetle (Pterostichus pensylvanicus)
  • Lianoe → (7 species)
    • Pterostichus macrogenys
    • Pterostichus macrocephalus
    • Pterostichus pseudopachinus
    • Pterostichus koheii
    • Pterostichus mirificus
    • Pterostichus mucronatus
    • Pterostichus sphodriformis
  • Pseudethira → (27 species)
    • Pterostichus balachowskyi
    • Pterostichus hoelli
    • Pterostichus perhoplites
    • Pterostichus tulobalu
    • Pterostichus dhorpatanicus
    • Pterostichus pseudoharmandi
    • Pterostichus fritzhiekei
    • Pterostichus ganja
    • Pterostichus letensis
    • Pterostichus conaensis
    • Pterostichus nepalensis
    • Pterostichus deuvesianus
    • Pterostichus hartmanni
    • Pterostichus olafi
    • Pterostichus deuvei
    • Pterostichus gagates
    • Pterostichus matsumurai
    • Pterostichus balu
    • Pterostichus immarginatus
    • Pterostichus gerdi
    • Pterostichus kleinfeldi
    • Pterostichus juga
    • Pterostichus janbritoi
    • Pterostichus angoarnigi
    • Pterostichus ganesh
    • Pterostichus chainapaani
    • Pterostichus gompanus
  • Platysma → (2 species)
    • Pterostichus leptis
    • Pterostichus niger
  • Melanius → (3 species)
    • Ground beetle (Pterostichus corvinus)
    • Pterostichus noguchii
    • Ground beetle (Pterostichus aterrimus)
  • Argutor → (7 species)
    • Argutor antiquus
    • Argutor pliocenicus
    • Pterostichus dulcis
    • A Ground Beetle (Pterostichus praetermissus)
    • Ground beetle (Pterostichus commutabilis)
    • A Ground Beetle (Pterostichus vernalis)
    • Pterostichus sulcitarsis
  • Steropus → (8 species)
    • Pterostichus alacer
    • Black Clock Beetle (Pterostichus madidus)
    • Pterostichus fasciatopunctatus
    • Steropes pici
    • Steropes obscurans
    • Steropes caspius
    • Steropes popei
    • Steropes latifrons
  • Petrophilus → (36 species)
    • A Ground Beetle (Pterostichus lachrymosus)
    • Pterostichus pertinax
    • A Beetle (Pterostichus coracinus)
    • Prosopogmus interstitialis
    • Prosopogmus suspectus
    • Prosopogmus punctifer
    • Prosopogmus yarrensis
    • Prosopogmus boisduvalii
    • Prosopogmus nitidipennis
    • Prosopogmus garavagliae
    • Prosopogmus sedlacekorum
    • Prosopogmus sulcatulus
    • Prosopogmus insperatus
    • Prosopogmus koghisensis
    • Prosopogmus austrinus
    • Prosopogmus namoyensis
    • Prosopogmus rubricornis
    • Prosopogmus opacidermis
    • Prosopogmus aoupiniensis
    • Prosopogmus batjanicus
    • Prosopogmus foveipennis
    • Prosopogmus hornabrooki
    • Prosopogmus occidentalis
    • Prosopogmus fortis
    • Prosopogmus irideus
    • Prosopogmus speculiferus
    • Prosopogmus harpaloides
    • Prosopogmus lescheni
    • Prosopogmus delicatulus
    • Prosopogmus oodiformis
    • Prosopogmus tasmanicus
    • Prosopogmus reichei
    • Prosopogmus savesi
    • Prosopogmus leai
    • Prosopogmus impressifrons
    • Prosopogmus monochrous
  • Gastrosticta → (2 species)
    • Pterostichus tumescens
    • Gastrosticta puta
  • Cryobius → (16 species)
    • Cryobius blanduloides
    • Cryobius delicatus
    • Pterostichus kurosawai
    • A Ground Beetle (Pterostichus brevicornis)
    • A Ground Beetle (Pterostichus riparius)
    • Pterostichus pumilio
    • A Ground Beetle (Pterostichus barryorum)
    • Pterostichus nivalis
    • A Ground Beetle (Pterostichus caribou)
    • A Ground Beetle (Pterostichus empetricola)
    • Pterostichus subsinuatus
    • Pterostichus nemoralis
    • A Ground Beetle (Pterostichus ventricosus)
    • Pterostichus negligens
    • Pterostichus pinguedineus
    • A Ground Beetle (Pterostichus surgens)
  • Pterostichus saxicola
  • Pterostichus ventralis
  • Pterostichus amurensis
  • Pterostichus ming
  • Pterostichus deceptrix
  • Pterostichus arribasi
  • Pterostichus zoiai
  • Pterostichus yoshikawai
  • Pterostichus gravis
  • Pterostichus dissimilis
  • Pterostichus nowitzkii
  • Pterostichus stictopleurus
  • Pterostichus kimurai
  • Pterostichus depressidorsis
  • Pterostichus liciniformis
  • Pterostichus dentellus
  • Pterostichus amblypterus
  • Pterostichus aristochroides
  • Pterostichus makalu
  • Pterostichus ochoticus
  • Pterostichus lumensis
  • Pterostichus pseudobarbarae
  • Pterostichus meisteri
  • Pterostichus merkli
  • Pterostichus arcuaticarinatus
  • Pterostichus walcotti
  • Pterostichus uchiyamai
  • Pterostichus dux
  • Pterostichus walteri
  • Pterostichus tokmakovae
  • Pterostichus schoenmanni
  • Pterostichus symmetricus
  • Pterostichus middendorffi
  • Pterostichus microbus
  • Pterostichus coruscus
  • Pterostichus transversalis
  • Pterostichus medvedevi
  • Pterostichus cantabricus
  • Pterostichus spathifer
  • Pterostichus caucasicola
  • Pterostichus krulikovskyi
  • Pterostichus naokii
  • Pterostichus ohsawacavus
  • Pterostichus sojaki
  • Pterostichus pasanauricus
  • Pterostichus malkini
  • Pterostichus davidi
  • Pterostichus unctulatus
  • Pterostichus quelpartensis
  • Pterostichus yamauchii
  • Pterostichus iwakiensis
  • Pterostichus kuwolsanensis
  • Pterostichus satyrus
  • Pterostichus marginellus
  • Pterostichus seticeps
  • A Ground Beetle (Pterostichus relictus)
  • Pterostichus mariamae
  • Pterostichus larisae
  • Pterostichus abishirensis
  • Pterostichus cursor
  • Pterostichus starcki
  • Pterostichus shinbodakensis
  • Pterostichus tenuimarginatus
  • Pterostichus pohnerti
  • Pterostichus yokohamae
  • Pterostichus takahashii
  • Pterostichus laticollis
  • Pterostichus justusii
  • Pterostichus oreophilus
  • Pterostichus nikodymi
  • Pterostichus ishizukai
  • Pterostichus bythiniensis
  • Pterostichus ruffoi
  • Pterostichus svetlanae
  • Pterostichus aksekianus
  • Pterostichus subgibbus
  • Pterostichus cantalicus
  • Pterostichus latemarginatus
  • Pterostichus basilobatus
  • Pterostichus kolymensis
  • Pterostichus pseudopercus
  • Pterostichus ompoensis
  • Pterostichus kalabi
  • Pterostichus yorikoae
  • Pterostichus pumpellyi
  • Pterostichus paulini
  • Pterostichus ulubeyensis
  • Pterostichus trinarius
  • Pterostichus shimizui
  • Pterostichus galaecianus
  • Pterostichus mosaicus
  • Pterostichus jogaesanensis
  • Pterostichus pseudoplatyderus
  • Pterostichus basipunctatus
  • Pterostichus jaechi
  • Pterostichus esakii
  • Pterostichus miyamai
  • Pterostichus casaleianus
  • Pterostichus compressus
  • Pterostichus strasseri
  • Pterostichus cordatissimus
  • Pterostichus watanabei
  • Pterostichus berezowskii
  • Pterostichus perlutus
  • Pterostichus sotkaensis
  • Pterostichus meyeri
  • Pterostichus curvatus
  • Pterostichus jani
  • Pterostichus pseudopilifer
  • Pterostichus glaferi
  • A Ground Beetle (Pterostichus parasimilis)
  • Pterostichus planctus
  • Pterostichus caucasicus
  • Pterostichus infimus
  • Pterostichus argutoriformis
  • Pterostichus irideus
  • Pterostichus triseriatus
  • Pterostichus acutipes
  • Pterostichus fornicatus
  • Pterostichus rufipes
  • Pterostichus planus
  • Pterostichus rugosipennis
  • Pterostichus peilingi
  • Pterostichus songoricus
  • Pterostichus depletus
  • Pterostichus cavicollis
  • Pterostichus talibani
  • Pterostichus glukhomanka
  • Pterostichus meskheticus
  • Pterostichus kaynashensis
  • Pterostichus zyzzovi
  • Pterostichus tatricus
  • Pterostichus jaechianus
  • Pterostichus pseudoplatysma
  • Pterostichus daihizanus
  • Pterostichus malaise
  • Pterostichus horstweiperti
  • Pterostichus duratii
  • Pterostichus muelleri
  • Pterostichus variabilis
  • Pterostichus focarilei
  • Pterostichus auriga
  • Pterostichus oblongiusculus
  • Pterostichus sagarmatha
  • Pterostichus gongga
  • Pterostichus devillei
  • Pterostichus subitus
  • Pterostichus audax
  • Pterostichus validus
  • Pterostichus acrogonus
  • Pterostichus provincialis
  • Pterostichus scitus
  • Pterostichus suffusus
  • Pterostichus fractus
  • Pterostichus labzuki
  • Pterostichus latistylis
  • Pterostichus chasautianus
  • Pterostichus nakanei
  • Pterostichus microps
  • Pterostichus daghestanus
  • Pterostichus bifoveolatus
  • Pterostichus sintanus
  • Pterostichus parviceps
  • Pterostichus bhutanensis
  • Pterostichus belizini
  • Pterostichus akkusianus
  • Pterostichus gurnetensis
  • Pterostichus sharanus
  • Pterostichus brancuccii
  • Pterostichus sejunctus
  • Pterostichus potanini
  • Pterostichus laborans
  • Pterostichus huashanus
  • Pterostichus liguricus
  • Pterostichus straneellus
  • Pterostichus tanakai
  • Pterostichus kopetzi
  • Pterostichus odontocnemis
  • Pterostichus balikliensis
  • Pterostichus pseudorotundus
  • Pterostichus korolkowi
  • Pterostichus marani
  • Pterostichus janoi
  • Pterostichus pilosus
  • Pterostichus filipjevi
  • Pterostichus kangchenjunga
  • Pterostichus tachongi
  • Pterostichus simillimus
  • Pterostichus vagepunctatus
  • Pterostichus necessarius
  • Pterostichus urengaicus
  • Pterostichus melanodes
  • Pterostichus septentrionis
  • Pterostichus mundus
  • Pterostichus leonisi
  • Pterostichus franzi
  • Pterostichus ussuriensis
  • Pterostichus krasnopolensis
  • Pterostichus kurikomasanus
  • Pterostichus setosicus
  • Pterostichus ishizuchiensis
  • Pterostichus selmanni
  • Pterostichus horvatovichi
  • Pterostichus chinlingshanicus
  • Pterostichus chogyesanus
  • Pterostichus bischoffianus
  • Pterostichus cylindriformis
  • Pterostichus rhanis
  • Pterostichus paulinoi
  • Pterostichus cucujinus
  • Pterostichus mattolenis
  • Pterostichus ordinarius
  • Pterostichus ordinatus
  • Pterostichus fukube
  • Pterostichus wenxianensis
  • Pterostichus lutshnikianus
  • Pterostichus gibbicollis
  • Pterostichus subiasi
  • Pterostichus septemtrionalis
  • Pterostichus kiangsu
  • Pterostichus variolatus
  • Pterostichus sejungendus
  • Pterostichus putus
  • Pterostichus starckianus
  • Pterostichus molopsoides
  • Pterostichus tienmushanus
  • Pterostichus korgei
  • Pterostichus obsidianus
  • Pterostichus phaeopus
  • Pterostichus teretis
  • Pterostichus nadari
  • Pterostichus chefsuricus
  • Pterostichus grajus
  • Pterostichus aralarensis
  • Pterostichus klapperichi
  • Pterostichus georgi
  • Pterostichus imurai
  • Pterostichus lucidus
  • Pterostichus phaeus
  • Pterostichus pantomus
  • Pterostichus crassiapex
  • Pterostichus anatolicus
  • Pterostichus ponticus
  • Pterostichus fencli
  • Pterostichus obesulus
  • Pterostichus latifianus
  • Pterostichus jureceki
  • Pterostichus nasuensis
  • Pterostichus avaricus
  • Pterostichus gobettii
  • Pterostichus orman
  • Pterostichus sasajii
  • Pterostichus faldermanni
  • Pterostichus bousqueti
  • Pterostichus chinkiangensis
  • Pterostichus iberlovi
  • Pterostichus cancellatus
  • Pterostichus findelii
  • Pterostichus sugimurai
  • Pterostichus monticoloides
  • Pterostichus quadraticollis
  • Pterostichus lacertus
  • Pterostichus himifuriho
  • Pterostichus pulcher
  • Pterostichus pseudopedius
  • A Ground Beetle (Pterostichus soperi)
  • Pterostichus ishii
  • Pterostichus yvanii
  • Pterostichus taradakensis
  • Pterostichus bifidiphallus
  • Pterostichus tamsii
  • Pterostichus ledenikensis
  • Pterostichus gotoensis
  • Pterostichus woronowi
  • Pterostichus bellatrix
  • Pterostichus smetanai
  • Pterostichus ophryoderus
  • Ground beetle (Pterostichus rostratus)
  • Pterostichus lutschniki
  • Pterostichus uralensis
  • Pterostichus suruganus
  • Pterostichus shirakamisanus
  • Pterostichus jugicola
  • Pterostichus goschi
  • Pterostichus altaiensis
  • Pterostichus consanguineus
  • Pterostichus taebaegsanus
  • Pterostichus gaoligongensis
  • Pterostichus costatus
  • Pterostichus alexeji
  • Pterostichus shulli
  • Pterostichus funakoshii
  • Pterostichus tosanus
  • Pterostichus orduensis
  • Pterostichus mellyi
  • Pterostichus acutidens
  • Pterostichus yulongshanensis
  • Pterostichus cunibakus
  • Pterostichus nivicola
  • Pterostichus decastriensis
  • Pterostichus brittoni
  • Pterostichus abrogatus
  • Pterostichus aapsorum
  • A Ground Beetle (Pterostichus ebeninus)
  • Pterostichus maderi
  • Pterostichus goriensis
  • Pterostichus megaloderus
  • Pterostichus creper
  • Pterostichus magoides
  • Pterostichus castanipes
  • Pterostichus setipes
  • Pterostichus procephalus
  • Pterostichus megacephalus
  • Pterostichus biocryus
  • Pterostichus ohbayashii
  • Pterostichus burmeisteri
  • Pterostichus fernquisti
  • Pterostichus shotaroi
  • Pterostichus yahikosanus
  • Pterostichus barbarae
  • Pterostichus dolens
  • Pterostichus carinatus
  • Pterostichus ikukoae
  • Pterostichus eriwanicus
  • Pterostichus masumotoi
  • Pterostichus napaea
  • Pterostichus parnassius
  • A Ground Beetle (Pterostichus oregonus)
  • Pterostichus rufitarsis
  • Pterostichus lingshanus
  • Pterostichus jacobsoni
  • Pterostichus shirozui
  • Pterostichus shilenkovi
  • Pterostichus insidiator
  • Pterostichus inopinatus
  • Pterostichus asturicus
  • Pterostichus montigena
  • Pterostichus tateishiyamanus
  • Pterostichus procerulus
  • Pterostichus eoyoritomus
  • Pterostichus latiusculus
  • Pterostichus malaisei
  • Pterostichus takadateyamanus
  • Pterostichus sachtlebeni
  • Pterostichus kiritshenkoi
  • Pterostichus jankowskyi
  • Pterostichus violaceotinctus
  • Pterostichus aubryi
  • Pterostichus jelepus
  • Pterostichus hirobane
  • Pterostichus commixtiformis
  • Pterostichus novotnyorum
  • Pterostichus yoshizawai
  • Pterostichus rutenus
  • Pterostichus properans
  • Pterostichus farkaci
  • Pterostichus palgongsanus
  • Pterostichus dufourii
  • Pterostichus pfizenmayeri
  • Pterostichus kleinfeldianus
  • Pterostichus arrowianus
  • Pterostichus schuelkei
  • Pterostichus cristicaudis
  • Pterostichus gliscans
  • Pterostichus chydaeus
  • Pterostichus uedaorum
  • Pterostichus dostali
  • Pterostichus pavani
  • Pterostichus variicornis
  • Pterostichus lugi
  • Pterostichus andreinii
  • Pterostichus geberti
  • Pterostichus lolo
  • Pterostichus toledanoi
  • Pterostichus epiroticus
  • Pterostichus capito
  • Pterostichus lacunosus
  • Pterostichus exilis
  • A Ground Beetle (Pterostichus mancus)
  • Pterostichus minutulus
  • Pterostichus tomensis
  • Pterostichus sponsor
  • Pterostichus miroshnikovi
  • Johnsons Waterfall Carabid Beetle (Pterostichus johnsoni)
  • Pterostichus champenoisi
  • Pterostichus mutoides
  • Pterostichus mannerheimii
  • Pterostichus foveolatus
  • Pterostichus illustris
  • Pterostichus consors
  • Pterostichus ainus
  • Pterostichus cordifer
  • Pterostichus itoi
  • Pterostichus oblongoparallelus
  • Pterostichus melas
  • Pterostichus rugosiceps
  • Pterostichus cratocephalus
  • Pterostichus eximius
  • Pterostichus giresuni
  • Pterostichus nishiyamai
  • A Ground Beetle (Pterostichus circulosus)
  • Pterostichus drescoi
  • Pterostichus rubripalpis
  • Pterostichus bullatus
  • Pterostichus laevigatus
  • Pterostichus emei
  • Pterostichus chujoi
  • Ground beetle (Pterostichus stygicus)
  • Pterostichus borcka
  • Pterostichus longiusculus
  • Pterostichus apfelbecki
  • Pterostichus svanicus
  • Pterostichus skoupyi
  • Pterostichus orionis
  • Pterostichus seriatus
  • Pterostichus virescens
  • Pterostichus enasanus
  • Pterostichus piceolus
  • Pterostichus montanus
  • Pterostichus spornyi
  • Pterostichus lukjanovitshi
  • Pterostichus belousovi
  • Pterostichus denticaudis
  • Pterostichus kucerai
  • Pterostichus punctiventris
  • Pterostichus mewakholensis
  • Pterostichus maurusiacus
  • Pterostichus platyops
  • Pterostichus turcomannicus
  • Pterostichus espagnoli
  • Pterostichus arcanoides
  • Pterostichus sakagamii
  • A Carabid Beetle (Pterostichus palmi)
  • Pterostichus aibgensis
  • Pterostichus ohkawai
  • Pterostichus atrox
  • Pterostichus andrewesi
  • Pterostichus kosakai
  • Pterostichus galae
  • Pterostichus hungaricus
  • Pterostichus omogoensis
  • Pterostichus tuberculiger
  • Pterostichus kurosai
  • Pterostichus quadriimpressus
  • Pterostichus cordaticollis
  • Pterostichus shirakamisan
  • Pterostichus xilingensis
  • Pterostichus reissi
  • Pterostichus blandulus
  • Pterostichus rudestriatus
  • Pterostichus aenescens
  • Pterostichus elmbergi
  • Pterostichus yamajii
  • Pterostichus styx
  • Pterostichus panticulatus
  • Pterostichus jugivagus
  • Pterostichus levadensis
  • Pterostichus ikizderensis
  • Pterostichus bituberculatus
  • Pterostichus schaschli
  • Pterostichus micans
  • Pterostichus scalptus
  • Pterostichus geojensis
  • Pterostichus magus
  • Pterostichus breviusculus
  • Pterostichus protensiformis
  • Pterostichus eboshiyamanus
  • Pterostichus leviculus
  • Pterostichus ogaensis
  • Pterostichus haftorni
  • Pterostichus euxinus
  • Pterostichus tropicalis
  • Pterostichus faunus
  • Pterostichus cervenkai
  • Pterostichus rifensis
  • Pterostichus chameleon
  • Pterostichus xatartii
  • Pterostichus karateghinicus
  • Pterostichus falcispinus
  • Pterostichus homalonotus
  • Pterostichus iripennis
  • Pterostichus raraensis
  • Pterostichus tokui
  • Pterostichus subabacipennis
  • Pterostichus safonovae
  • Pterostichus comatus
  • Pterostichus innshanensis
  • Pterostichus lodosi
  • Pterostichus dudkoi
  • Pterostichus kongosanus
  • Pterostichus eobius
  • Pterostichus giacomazzoi
  • Pterostichus clepsydra
  • Pterostichus zhejiangensis
  • Pterostichus ehlersi
  • Pterostichus przewalskyi
  • Pterostichus abnormis
  • Pterostichus alexandrovi
  • Pterostichus mundatus
  • Pterostichus exceptus
  • Pterostichus malissorum
  • Pterostichus kadjaranze
  • Pterostichus trubilini
  • Pterostichus hisamatsui
  • Pterostichus kerzhneri
  • Pterostichus sterbai
  • Pterostichus phungaraziensis
  • Pterostichus kadleci
  • Pterostichus espanoli
  • Pterostichus koenigianus
  • Pterostichus confucius
  • Pterostichus filum
  • Pterostichus bowanus
  • Pterostichus reuteri
  • Pterostichus rotundus
  • Pterostichus ferreroi
  • A Ground Beetle (Pterostichus atratus)
  • Pterostichus porcellus
  • Pterostichus catei
  • Pterostichus macedonicus
  • Pterostichus hejkali
  • Pterostichus coreicus
  • Pterostichus ripensis
  • Pterostichus funestes
  • Pterostichus shingarevi
  • Pterostichus strigosus
  • Pterostichus capitolinus
  • Pterostichus carri
  • Pterostichus strigipennis
  • Pterostichus ybousqueti
  • Pterostichus ixion
  • Pterostichus hirasawai
  • Pterostichus bisulcitarsis
  • Pterostichus cacumensis
  • Pterostichus cecchiniae
  • Pterostichus triangularis
  • Pterostichus mediocris
  • Pterostichus batesi
  • Pterostichus multinodosus
  • Pterostichus neilgaimani
  • Ground beetle (Pterostichus caudicalis)
  • Pterostichus incommodus
  • Pterostichus ovaliphallus
  • Pterostichus wrasei
  • Pterostichus abudarini
  • Pterostichus migliaccioi
  • Pterostichus adatarasanus
  • Pterostichus enodis
  • Pterostichus jaljaleensis
  • Pterostichus ignavus
  • Pterostichus depilatus
  • Pterostichus abasinus
  • Pterostichus acuspina
  • Pterostichus punctibasalis
  • Pterostichus casalei
  • Pterostichus colasi
  • Pterostichus aemiliae
  • Pterostichus santostamangi
  • Pterostichus arsenjevi
  • Pterostichus drescheri
  • Pterostichus satoi
  • Pterostichus leica
  • Pterostichus ferghanicus
  • Pterostichus pseudodiversus
  • Pterostichus kambaiti
  • Pterostichus thorectoides
  • Pterostichus komalus
  • Pterostichus masahiroi
  • Pterostichus odvarkai
  • Pterostichus dandongensis
  • Pterostichus altaianus
  • Pterostichus dentifer
  • Pterostichus orobius
  • Pterostichus michailovi
  • Pterostichus makolskii
  • Pterostichus barlensis
  • Pterostichus gallega
  • Pterostichus militaris
  • Pterostichus rufonitens
  • Pterostichus orion
  • Pterostichus piliferus
  • Pterostichus komiyai
  • Pterostichus curtatus
  • Pterostichus jakobsonianus
  • Pterostichus tanjae
  • Pterostichus planicola
  • Pterostichus laferi
  • Pterostichus sutschanensis
  • Pterostichus crenulatopunctatus
  • Pterostichus tantillus
  • Pterostichus analis
  • Pterostichus takaosanus
  • Pterostichus beneshi
  • Pterostichus oenotrius
  • Pterostichus gerstlensis
  • Pterostichus glabripennis
  • Pterostichus mitoyamanus
  • A Ground Beetle (Pterostichus vermiculosus)
  • Pterostichus procax
  • Pterostichus kokeilii
  • Pterostichus bruckii
  • Pterostichus poppiusi
  • Pterostichus muehlfeldii
  • Pterostichus blumenthali
  • Pterostichus tamarae
  • Pterostichus bureli
  • Pterostichus hiramatsui
  • Pterostichus hoplites
  • Pterostichus barbarinus
  • Pterostichus amanoi
  • Pterostichus bungei
  • Pterostichus diversus
  • Pterostichus morawitzianus
  • Pterostichus nicaeensis
  • Pterostichus kurentzovi
  • Pterostichus rugiceps
  • Pterostichus schoenherri
  • Pterostichus calvitarsis
  • Pterostichus armenus
  • A Ground Beetle (Pterostichus novus)
  • Pterostichus monostigma
  • Pterostichus externepunctatus
  • Pterostichus percontator
  • Pterostichus altaicus
  • Pterostichus woongbii
  • Pterostichus thorectes
  • Pterostichus schrettenbrunneri
  • Pterostichus tsurugiyamanus
  • Pterostichus hubeicus
  • Pterostichus fujisanus
  • A Ground Beetle (Pterostichus arcticola)
  • Pterostichus fulvescens
  • Pterostichus herzi
  • Pterostichus szetschuanensis
  • Pterostichus nigripalpis
  • Pterostichus shojii
  • Pterostichus hozumii
  • Pterostichus vladivostokensis
  • Ground beetle (Pterostichus corrusculus)
  • Pterostichus lasserrei
  • Pterostichus maximus
  • Pterostichus rugosus
  • Pterostichus mujahedeeni
  • Pterostichus toyodai
  • Pterostichus theeli
  • Pterostichus auratus
  • A Ground Beetle (Pterostichus primus)
  • Pterostichus agilis
  • Pterostichus tsukubasanus
  • Pterostichus lassulus
  • Pterostichus restrictus
  • Pterostichus rasilis
  • Pterostichus variseta
  • Pterostichus punctiger
  • Pterostichus eschscholtzii
  • Pterostichus esuriens
  • Pterostichus sakuraii
  • Pterostichus gromykoi
  • Pterostichus tichomirovi
  • Pterostichus reiseri
  • Pterostichus chokaisanus
  • Pterostichus tonkinensis
  • Pterostichus okiensis
  • Pterostichus semirugosus
  • Pterostichus olegi
  • A Ground Beetle (Pterostichus chipewyan)
  • Pterostichus ishiharai
  • Pterostichus muellermotzfeldi
  • Pterostichus burkhan
  • Pterostichus akitai
  • Pterostichus globosus
  • Pterostichus longipennis
  • Pterostichus yamizosanus
  • Pterostichus kadoudali
  • Pterostichus jiricola
  • Pterostichus lombardus
  • Pterostichus imitatus
  • Pterostichus kalo
  • Pterostichus amadeus
  • Pterostichus mirus
  • Pterostichus keltiekus
  • Pterostichus petulans
  • Pterostichus montisdeorum
  • Pterostichus dilutipes
  • Pterostichus pseudoastutus
  • Pterostichus schmidtii
  • Pterostichus ademidovae
  • Pterostichus tottoriensis
  • Pterostichus subaeneus
  • Pterostichus scutellaris
  • Pterostichus mariae
  • Pterostichus nimbatidius
  • Pterostichus microcephalus
  • Pterostichus viridellus
  • Pterostichus jurecekianus
  • Pterostichus catalonicus
  • Pterostichus mizunoyai
  • Pterostichus pergracilis
  • Pterostichus dundai
  • Pterostichus woodi
  • Pterostichus abensis
  • Pterostichus sediorcus
  • Pterostichus pusillus
  • Pterostichus singularis
  • Pterostichus licenti
  • Pterostichus brevis
  • Pterostichus harponifer
  • Pterostichus aimaki
  • Pterostichus colonus
  • Pterostichus machulkai
  • Pterostichus turanensis
  • Pterostichus patruelis
  • Pterostichus palekhai
  • Pterostichus distinctus
  • A Ground Beetle (Pterostichus kotzebuei)
  • Pterostichus craterensis
  • Pterostichus nishidai
  • Pterostichus okutamae
  • Pterostichus humidulus
  • Pterostichus dauricus
  • Pterostichus tukchensis
  • Pterostichus macrothorax
  • Pterostichus datshenkoae
  • Pterostichus asahinus
  • Pterostichus ishikawai
  • Pterostichus ohkurai
  • Pterostichus janatai
  • Pterostichus pithyusicus
  • Pterostichus musashiensis
  • Pterostichus validior
  • Pterostichus chechcirensis
  • Pterostichus sayanus
  • Pterostichus aereipennis
  • Pterostichus kolosovi
  • Pterostichus distinctissimus
  • A Carabid Beetle (Pterostichus lubricus)
  • Pterostichus hikosanus
  • Pterostichus bhadravati
  • Pterostichus vecors
  • Pterostichus rousi
  • Pterostichus pulchellus
  • Pterostichus arcanus
  • Pterostichus impressicollis
  • Pterostichus tuberifer
  • Pterostichus ssemenovi
  • Pterostichus annosus
  • Pterostichus mukdenensis
  • Pterostichus kultianus
  • Pterostichus baldwini
  • Pterostichus maichensis
  • Pterostichus destitutus
  • Pterostichus kaniei
  • Pterostichus lanista
  • Pterostichus campbelli
  • Pterostichus abagonensis
  • Pterostichus chenpengi
  • Pterostichus ghilarovi
  • Pterostichus tatianae
  • Pterostichus fenestratus
  • Pterostichus haesitatus
  • Pterostichus kirschenblatti
  • Pterostichus longipes
  • Pterostichus jungens
  • Pterostichus amagisanus
  • Pterostichus arnoldii
  • Pterostichus rhilensis
  • Pterostichus fujimurai
  • A Ground Beetle (Pterostichus carolinus)
  • Pterostichus matveichuki
  • Pterostichus sinicus
  • Pterostichus anomostriatus
  • A Carabid Beetle (Pterostichus hypogeus)
  • Pterostichus imasakai
  • Pterostichus kitakamisanus
  • Pterostichus schodaicus
  • Pterostichus shennongjianus
  • Pterostichus brevipennis
  • Pterostichus setosus
  • Pterostichus cribratus
  • Pterostichus expeditus
  • Pterostichus weigeli
  • Pterostichus laevipunctatus
  • Pterostichus vseteckai
  • Pterostichus sudrei
  • Pterostichus aanistschenkoi
  • Pterostichus heinzianus
  • Pterostichus abaxoides
  • Pterostichus amoenus
  • Pterostichus tanakaorum
  • Pterostichus ishikawaioides
  • Pterostichus masatakai
  • Pterostichus tundrae
  • Pterostichus heptapotamicus
  • Pterostichus gwervaenus
  • Pterostichus occultus
  • Pterostichus arator
  • Pterostichus satunini
  • Pterostichus fallettii
  • Pterostichus nodicornis
  • Pterostichus carsticus
  • Pterostichus hedkeinekus
  • Pterostichus vandykei
  • Pterostichus baenningeri
  • Pterostichus adulterinus
  • Pterostichus mulensis
  • Pterostichus heilongjiangensis
  • Pterostichus perisi
  • Pterostichus stomoides
  • Pterostichus sierranus
  • Pterostichus krali
  • Pterostichus zolotarewi
  • Pterostichus ienishteai
  • Pterostichus akozyrevi
  • Pterostichus kaninensis
  • Pterostichus gyrosus
  • Pterostichus rousselli
  • Pterostichus latitemporis
  • Pterostichus ferreri
  • Pterostichus insignicollis
  • Pterostichus tsurugaensis
  • Pterostichus kamtschaticus
  • Pterostichus exochus
  • Pterostichus ingusha
  • Pterostichus kvirensis
  • Pterostichus satsumanus
  • Pterostichus peninsularis
  • Pterostichus yunnanensis
  • Pterostichus besucheti
  • Pterostichus castor
  • Pterostichus lamuticus
  • Pterostichus subtilissimus
  • Pterostichus sumondakensis
  • Pterostichus isumiensis
  • Pterostichus lesticoides
  • Pterostichus sodalicius
  • Pterostichus brevilama
  • Pterostichus championi
  • Pterostichus sudai
  • Pterostichus galinae
  • Pterostichus tiliaceoradix
  • Pterostichus swaneticus
  • Pterostichus sungariensis
  • Pterostichus ovalipennis
  • Pterostichus saueri
  • Pterostichus harmandi
  • Pterostichus yoshidai
  • Pterostichus hakusanus
  • Pterostichus deino
  • A Ground Beetle (Pterostichus bryanti)
  • Pterostichus glacialis
  • Pterostichus heinzi
  • Pterostichus ghilianii
  • Pterostichus cathaicus
  • Pterostichus cylindricus
  • Pterostichus matchai
  • Pterostichus bryantoides
  • Pterostichus andreae
  • Pterostichus balthasari
  • Pterostichus tenenbaumianus
  • Pterostichus tokejii
  • Pterostichus illigeri
  • Pterostichus poppiusianus
  • Pterostichus honnoratii
  • Pterostichus barri
  • Pterostichus chotanensis
  • Pterostichus isolatus
  • Pterostichus punctisternus
  • Pterostichus pseudostuxbergi
  • Pterostichus mascareauxi
  • Pterostichus regeli
  • Pterostichus pedemontanus
  • Pterostichus buglaniensis
  • Pterostichus orestes
  • Pterostichus diligendus
  • Pterostichus mercedianus
  • Pterostichus yatsuensis
  • Pterostichus bhunetansis
  • Pterostichus dormitans
  • Pterostichus destructus
  • Pterostichus molestus
  • Pterostichus ryomoensis
  • Pterostichus pentheri
  • Pterostichus scuticollis
  • Pterostichus spraguei
  • Pterostichus doris
  • Pterostichus ottomanus
  • Pterostichus wittmeri
  • Pterostichus tschitscherinianus
  • Pterostichus weiperti
  • A Ground Beetle (Pterostichus tareumiut)
  • Pterostichus liodactylus
  • Pterostichus stoeckleini
  • Pterostichus miroslavi
  • Pterostichus nordqvisti
  • Pterostichus seungmoi
  • Pterostichus gassanus
  • Pterostichus kiiensis
  • A Ground Beetle (Pterostichus lustrans)
  • Pterostichus nigellatus
  • A Ground Beetle (Pterostichus femoralis)
  • Pterostichus solskyi
  • Pterostichus semenowi
  • Pterostichus cryobioides
  • Pterostichus araraticus
  • Pterostichus microphthalmus
  • Pterostichus shaovuensis
  • Pterostichus rengensis
  • Pterostichus falli
  • Pterostichus stantonensis
  • Pterostichus pratti
  • Pterostichus superciliosus
  • Pterostichus macer
  • Pterostichus minax
  • A Ground Beetle (Pterostichus similis)
  • Pterostichus astutus
  • Pterostichus troglophilus
  • Pterostichus darvazicus
  • Pterostichus nasui
  • Pterostichus hidanus
  • Pterostichus zamotajlovi
  • Pterostichus arrowi
  • Pterostichus dipojranus
  • Pterostichus capitatus
  • Pterostichus mandzhuricus
  • Pterostichus ohdaisanus
  • Pterostichus bielzii
  • Pterostichus tahirai
  • Pterostichus szekessyianus
  • Pterostichus mandibularoides
  • Pterostichus glabricollis
  • Pterostichus rolex
  • Pterostichus validiceps
  • Pterostichus parvicollis
  • Pterostichus sinensis
  • Pterostichus apiculatiphallus
  • A Ground Beetle (Pterostichus hudsonicus)
  • Pterostichus sacheri
  • Pterostichus subacutus
  • Pterostichus rousianus
  • Pterostichus dorjulensis
  • Pterostichus noburai
  • Pterostichus impressus
  • Pterostichus pachinus
  • Pterostichus colchicus
  • Pterostichus vicinus
  • Pterostichus rufopiceus
  • Pterostichus todai
  • Pterostichus planiusculus
  • Pterostichus annamita
  • Pterostichus gallopavo
  • Pterostichus planaticollis
  • Pterostichus insidatrix
  • Pterostichus lineatopunctatus
  • Pterostichus modicellus
  • Pterostichus vetustus
  • Pterostichus hornii
  • Pterostichus asiaticus
  • Pterostichus amaroides
  • Pterostichus neglectus
  • Pterostichus shavrini
  • Pterostichus thanglaensis
  • Pterostichus firmus
  • Pterostichus syleus
  • Pterostichus discrepans
  • Pterostichus perlamatus
  • Pterostichus platyderus
  • Pterostichus cavazzutii
  • Pterostichus guizhouensis
  • Pterostichus chungkingi
  • Pterostichus kalhys
  • Pterostichus miehei
  • Pterostichus kataevi
  • Pterostichus lidarus
  • Pterostichus apenninus
  • Pterostichus comorus
  • Pterostichus elongatus
  • Pterostichus yanoi
  • Pterostichus defossus
  • Pterostichus josephi
  • Pterostichus stricticollis
  • Pterostichus ambiguus
  • Pterostichus antepunctatus
  • Pterostichus schneideri
  • Pterostichus nigellus
  • Pterostichus crassiusculus
  • Pterostichus koenigi
  • Pterostichus staveni
  • Pterostichus maeklini

Other links

• Wikidata
• Wikipedia (language: en)
• Wikipedia (language: de)
• Wikipedia (language: es)
• Wikipedia (language: fr)
• Wikipedia (language: ru)
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• NCBI (genetics)"
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