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Base of the evolutionary tree for rove beetles (Staphylinoidea)

rove beetles (Staphylinoidea) (70668 species)
- unnamed group 1681231 → (68696 species)
- unnamed group 1679260 → (1972 species)
  - Minute Moss Beetle (Hydraenidae) → (1148 species)
  - featherwing beetles (Ptiliidae) → (824 species)
    - Acrotrichinae → (319 species)
      - Acrotrichis → (225 species)
      - Ptinella → (60 species)
      - Nephanes → (14 species)
      - Baeocrara → (7 species)
      - Smicrus → (13 species)
    - Nellosana → (5 species)
      - Nellosana insperata
      - Nellosana elegantula
      - Nellosana minima
      - Nellosana intermedia
      - Nellosana grandis
    - Actinopteryx → (7 species)
      - Actinopteryx reflexa
      - Actinopteryx tillyardi
      - Actinopteryx parallela
      - Actinopteryx hoguei
      - Actinopteryx hercules
      - Actinopteryx lancifer
      - Actinopteryx fucicola
    - Urotriainus → (3 species)
      - Urotriainus molesti
      - Urotriainus bidentatus
      - Urotriainus grandis
    - Millidium → (3 species)
      - Millidium minutissimum
      - Millidium sculpturatum
      - Millidium triramosum
    - Ptiliinae → (198 species)
      - Ptenidium → (73 species)
      - Nanosella → (7 species)
      - Micridium → (17 species)
      - Ptilium → (44 species)
      - Microptilium → (2 species)
      - Ptiliola → (5 species)
      - Ptiliolum → (23 species)
      - Cylindroselloides → (2 species)
      - Nossidium → (8 species)
      - Euryptilium → (2 species)
      - Pteryx → (11 species)
      - Baranowskiella ehnstromi
      - Scydosella musawasensis
      - Motschulskium sinuatocolle
      - Throscoptiloides norcoensis
    - Gomyella → (13 species)
      - Gomyella lata
      - Gomyella obscura
      - Gomyella parva
      - Gomyella simplex
      - Gomyella rufotestacea
      - Gomyella apteroides
      - Gomyella carinata
      - Gomyella similata
      - Gomyella lawrencei
      - Gomyella longicornis
      - Gomyella canaliculata
      - Gomyella aptera
      - Gomyella schauenbergi
    - Storicricha → (7 species)
      - Storicricha tumida
      - Storicricha furcilla
      - Storicricha primitiva
      - Storicricha furiosa
      - Storicricha nigrita
      - Storicricha magnifica
      - Storicricha copiosa
    - Sindosium → (2 species)
      - Sindosium foveatum
      - Sindosium opacipenne
    - Ussurilumpia → (2 species)
      - Ussurilumpia trichaptumi
      - Ussurilumpia trihaptumi
    - Actidium → (31 species)
      - Actidium politum
      - Actidium boudieri
      - Actidium variolatum
      - Actidium crotchianum
      - Actidium sharpianum
      - Actidium magnum
      - Actidium interpolatum
      - Actidium reticulatum
      - Actidium delicatulum
      - Actidium reitteri
      - Actidium pruinosum
      - Actidium kraatzi
      - Actidium coarctatum
      - Actidium fowlerianum
      - Actidium mascarenhasi
      - Actidium flavoterminum
      - Actidium lineare
      - Actidium leleupi
      - Actidium producta
      - Actidium longicorne
      - Actidium granulosum
      - Actidium cameroni
      - Actidium rotundicolle
      - Actidium truncatum
      - Actidium angulicolle
      - Actidium parvulum
      - Actidium attenuatum
      - Actidium aegypti
      - Actidium robustulum
      - Actidium aterrimum
      - Actidium difficilis
    - Limulopteryx → (2 species)
      - Limulopteryx hintelmanni
      - Limulopteryx loebli
    - Isolumpia → (2 species)
      - Isolumpia divina
      - Isolumpia propeflava
    - Africoptilium → (3 species)
      - Africoptilium concinnum
      - Africoptilium mimicum
      - Africoptilium marginatum
    - Ptinellodes → (8 species)
      - Ptinellodes lecontei
      - Ptinellodes heterosternus
      - Ptinellodes suteri
      - Ptinellodes similis
      - Ptinellodes aldabricus
      - Ptinellodes darwini
      - Ptinellodes hunteri
      - Ptinellodes malkini
    - Erro → (2 species)
      - Erro angolensis
      - Erro impiger
    - Bambara → (22 species)
      - Bambara brunnea
      - Bambara problematica
      - Bambara steevesi
      - Bambara contorta
      - Bambara dybasi
      - Bambara edwardi
      - Bambara invisibilis
      - Bambara merina
      - Bambara nidicola
      - Bambara thomasi
      - Bambara frosti
      - Bambara joannis
      - Bambara verecunda
      - Bambara sublutea
      - Bambara secubita
      - Bambara testacea
      - Bambara lanquidula
      - Bambara fusca
      - Bambara elongatula
      - Bambara intricata
      - Bambara wagneri
      - Bambara opaca
    - Notoptenidium → (12 species)
      - Notoptenidium subitum
      - Notoptenidium johnsoni
      - Notoptenidium sparsum
      - Notoptenidium lawsoni
      - Notoptenidium kuscheli
      - Notoptenidium parvum
      - Notoptenidium apterum
      - Notoptenidium flavum
      - Notoptenidium aubrooki
      - Notoptenidium similatum
      - Notoptenidium crassum
      - Notoptenidium oblongum
    - Fenestellidium → (3 species)
      - Fenestellidium kakamegaense
      - Fenestellidium canfordanum
      - Fenestellidium capense
    - Throscidium → (2 species)
      - Throscidium fairmairii
      - Throscidium germainii
    - Porophila → (4 species)
      - Porophila malkini
      - Porophila mystacea
      - Porophila cedri
      - Porophila lazovskii
    - Mikado → (4 species)
      - Mikado japonicus
      - Mikado agilis
      - Mikado parvicornis
      - Mikado parva
    - Oligella → (4 species)
      - Oligella intermedia
      - Oligella foveolata
      - Oligella nana
      - Oligella insignis
    - Phytotelmatrichis → (3 species)
      - Phytotelmatrichis rava
      - Phytotelmatrichis peruviensis
      - Phytotelmatrichis osopaddington
    - Bicavella → (2 species)
      - Bicavella nossidioides
      - Bicavella relicta
    - Ptiliodes → (13 species)
      - Ptiliodes purpurascens
      - Ptiliodes pruinosus
      - Ptiliodes pulchellus
      - Ptiliodes australis
      - Ptiliodes curtus
      - Ptiliodes austerus
      - Ptiliodes naufragus
      - Ptiliodes congener
      - Ptiliodes insignis
      - Ptiliodes posticalis
      - Ptiliodes blackburni
      - Ptiliodes amplicollis
      - Ptiliodes cavifrons
    - Myrmicotrichis → (2 species)
      - Myrmicotrichis aequatorialis
      - Myrmicotrichis subvittata
    - Dacrysoma → (17 species)
      - Dacrysoma imitatum
      - Dacrysoma ravelosoni
      - Dacrysoma fusum
      - Dacrysoma dolorosum
      - Dacrysoma varium
      - Dacrysoma rahanitriniainae
      - Dacrysoma felis
      - Dacrysoma denticulatum
      - Dacrysoma nigerrimum
      - Dacrysoma subinsulsum
      - Dacrysoma andasibense
      - Dacrysoma ranomafanense
      - Dacrysoma fabrum
      - Dacrysoma delicatum
      - Dacrysoma longulum
      - Dacrysoma usambarense
      - Dacrysoma pilosum
    - Discheramocephalus → (15 species)
      - Discheramocephalus elisabethae
      - Discheramocephalus interfusus
      - Discheramocephalus jarmilae
      - Discheramocephalus inretitus
      - Discheramocephalus parvus
      - Discheramocephalus mikaeli
      - Discheramocephalus brucei
      - Discheramocephalus malalae
      - Discheramocephalus stewarti
      - Discheramocephalus capac
      - Discheramocephalus minutissimus
      - Discheramocephalus vasilii
      - Discheramocephalus semisulcatus
      - Discheramocephalus bisulcatus
      - Discheramocephalus angustus
    - Rodwayia → (7 species)
      - Rodwayia orientalis
      - Rodwayia hirsuta
      - Rodwayia occidentalis
      - Rodwayia ovata
      - Rodwayia minuta
      - Rodwayia intercoxalis
      - Rodwayia grandis
    - Dipentium → (15 species)
      - Dipentium reticulatum
      - Dipentium neglectum
      - Dipentium castaneum
      - Dipentium zelandicum
      - Dipentium punctatum
      - Dipentium gossi
      - Dipentium japonicum
      - Dipentium constrictum
      - Dipentium latum
      - Dipentium parvum
      - Dipentium incertum
      - Dipentium wolffi
      - Dipentium parvicolle
      - Dipentium magnum
      - Dipentium amplicolle
    - Kuschelidium → (2 species)
      - Kuschelidium maori
      - Kuschelidium okinawense
    - horse-shoe crab beetles (Cephaloplectinae) → (21 species)
      - Limulodes → (15 species)
      - Cephaloplectus → (6 species)
    - Cissidium → (25 species)
      - Cissidium scutellare
      - Cissidium petri
      - Cissidium ishigakiense
      - Cissidium foveolatum
      - Cissidium globosum
      - Cissidium rufescens
      - Cissidium latum
      - Cissidium banari
      - Cissidium sawadai
      - Cissidium reitteri
      - Cissidium sakaii
      - Cissidium lisae
      - Cissidium nomurai
      - Cissidium matthewsi
      - Cissidium shibatai
      - Cissidium crowsoni
      - Cissidium nishikawai
      - Cissidium itoi
      - Cissidium amboroensis
      - Cissidium tigrum
      - Cissidium ikeuchii
      - Cissidium adustipenne
      - Cissidium elongatum
      - Cissidium basale
      - Cissidium okuense
    - Leaduadicus → (2 species)
      - Leaduadicus imperialis
      - Leaduadicus tolerabilis
    - Leptinla elongata
    - Championella nubigena
    - Chaska nawi
    - Throscoptilium duryi
    - Paralimulodes wasmanni
    - Fijisella vituensis
    - Neotrichopteryx grandelytra
    - Nepalumpia markusi
    - Astatopteryx laticollis
    - Eulimulodes mexicanus
    - Throscosana pecki
    - Pterycodes salvinii
    - Babrama arcana
    - Phililumpia werneri
    - Pycnopteryx schmidti
    - Hydnosella globitheca
    - Petrotrichis rotundata
    - Kekveus jason
    - Dybasina productus
    - Microtrichis pectana
    - Fijiselloides nandaivatuensis
    - Cylindrosella dampfi
    - Etronia convexa
    - Garicaphila pilosa
    - Malkinella cavatica
    - Limulosella waspucensis
    - Xenopteryx setosus
    - Ptenidotonium longicorne
    - Suterina microcephala
    - Ptiliopycna moerens
    - Seminis factiosum
    - Greensladella semipunctata
    - Cochliarion victoriense
    - Chirostirca gomyi
    - Primorskiella anodonta
    - Kimoda globosa
    - Tasmangarica chisholmi
    - Vitusella fijiensis
    - Sikhotelumpia dersuuzalai
    - Scydoselloides alajuelaensis
    - Nellosanoides thayerae
    - Paratuposa placentis

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