Base of the evolutionary tree for bombadier beetles (Brachininae)

← parent

• bombadier beetles (Brachininae) (701 species)
  • Brachinini → (591 species)
    • Bombardier Beetle (Brachinus) → (337 species)
      • Brachinus hylaenus
      • Brachinus subnotatus
      • Brachinus elegans
      • Brachinus cinctellus
      • Brachinus albarracinus
      • Brachinus hamatus
      • Brachinus adustipennis
      • Brachinus tsara
      • Brachinus dilatipennis
      • Brachinus lombokensis
      • A Ground Beetle (Brachinus cyanipennis)
      • Brachinus tetraspilotus
      • A Carabid Beetle (Brachinus patruelis)
      • Brachinus melancholicus
      • Brachinus andreaei
      • Brachinus promontorii
      • A Ground Beetle (Brachinus alternans)
      • Brachinus costiger
      • Brachinus sexmaculatus
      • Brachinus afghanus
      • Brachinus sallei
      • Brachinus marginellus
      • Brachinus armiger
      • Brachinus peltastes
      • Brachinus tenuis
      • Brachinus atripensis
      • Brachinus eucosmus
      • Brachinus psophia
      • Brachinus illotus
      • Brachinus melanarthrus
      • Brachinus pachygaster
      • Brachinus macrocerus
      • Brachinus distans
      • Brachinus punctaticollis
      • Brachinus servillei
      • Brachinus limbellus
      • Brachinus dryas
      • Brachinus fageli
      • Brachinus hazardi
      • Brachinus sexnotatus
      • Brachinus circumtinctus
      • Brachinus americanus
      • Brachinus plagiatus
      • Brachinus aabaaba
      • Brachinus mesopotamicus
      • Brachinus rugipennis
      • A Ground Beetle (Brachinus tenuicollis)
      • Brachinus cruciatus
      • Brachinus connectoides
      • Brachinus oaxacensis
      • Brachinus limbicollis
      • Brachinus daliensis
      • Brachinus suensoni
      • Brachinus papua
      • Brachinus talyschensis
      • Brachinus testaceus
      • Brachinus xanthophryus
      • Brachinus brunneus
      • Brachinus cambodgensis
      • Brachinus cychroides
      • Brachinus midoli
      • Brachinus jakli
      • Brachinus micheli
      • Brachinus fulvipennis
      • Brachinus limbiger
      • A Ground Beetle (Brachinus fulminatus)
      • Brachinus favicollis
      • Brachinus repressus
      • Brachinus mauretanicus
      • Brachinus hirsutus
      • Brachinus dorsalis
      • Brachinus diffusus
      • Brachinus ghindanus
      • Brachinus flavus
      • Brachinus mactus
      • Brachinus chalchihuitlicue
      • Brachinus niger
      • Brachinus lateralis
      • Brachinus magyari
      • Brachinus variventris
      • Brachinus rikatlae
      • Brachinus evanescens
      • Brachinus humeralis
      • Brachinus obtusus
      • Brachinus ichabodopsis
      • Brachinus nigridorsis
      • Brachinus minor
      • Brachinus gebhardis
      • Brachinus methneri
      • Brachinus cibolensis
      • Brachinus sicardi
      • Brachinus lesnei
      • Brachinus kollari
      • Brachinus dilottii
      • Brachinus microrrhabdus
      • Brachinus bantimurungensis
      • Brachinus maublanci
      • Brachinus imporcitis
      • Brachinus alexandri
      • Brachinus suturellus
      • Brachinus collarti
      • Brachinus atramentarius
      • Streaked Bombardier (Brachinus sclopeta)
      • Brachinus madecassus
      • Brachinus langenhani
      • Brachinus fukienensis
      • Brachinus nigricans
      • Brachinus sonorous
      • Brachinus luzonicus
      • Brachinus frontalis
      • Brachinus nigrovirescens
      • Brachinus quadripennis
      • Brachinus puncticollis
      • Brachinus boeticus
      • Brachinus tianshanicus
      • Brachinus costulipennis
      • Brachinus newberryi
      • Brachinus sublaevis
      • Brachinus turnai
      • Brachinus proximus
      • Brachinus leprieuri
      • Brachinus alexiguus
      • Brachinus lethierryi
      • Brachinus palawanensis
      • Brachinus aeneicostis
      • Brachinus demoulini
      • Brachinus flavipes
      • Brachinus abbreviatus
      • Brachinus constrictus
      • Brachinus perplexus
      • Brachinus vulcanoides
      • Brachinus bilineatus
      • Brachinus reyi
      • Brachinus brittoni
      • Brachinus pictus
      • Brachinus turkestanicus
      • Brachinus hubeiensis
      • Brachinus mobilis
      • Brachinus microamericanus
      • Brachinus agraphus
      • Brachinus bayardi
      • Brachinus dilatatus
      • Brachinus velutinus
      • Brachinus azureipennis
      • Brachinus nigricornis
      • Brachinus truncatulus
      • Brachinus elegantulus
      • Brachinus phaeocerus
      • Brachinus tetrastigma
      • Brachinus imperialensis
      • Brachinus scriptus
      • Brachinus laetus
      • Brachinus kansanus
      • Brachinus brevicollis
      • Brachinus cinctipennis
      • Brachinus costatulus
      • Brachinus simulans
      • Brachinus stenoderus
      • Brachinus costipennis
      • Brachinus merkli
      • Brachinus arboreus
      • Brachinus immarginatus
      • Brachinus bruchi
      • Brachinus dalatensis
      • Brachinus solidus
      • Brachinus stappersi
      • Brachinus geniculatus
      • Brachinus batuensis
      • Brachinus caffer
      • Brachinus chinensis
      • Brachinus paviei
      • Brachinus suturalis
      • Brachinus viridipennis
      • Brachinus elongatulus
      • Brachinus exquisitus
      • Brachinus texanus
      • Brachinus stygius
      • Brachinus perrieri
      • Brachinus conformis
      • Brachinus longipalpis
      • A Carabid Beetle (Brachinus ovipennis)
      • Brachinus olgae
      • Brachinus andalusiacus
      • Brachinus galactoderus
      • Brachinus capnicus
      • Brachinus catalonicus
      • Brachinus oblongus
      • Brachinus genicularis
      • Brachinus somereni
      • Brachinus aristokrates
      • Brachinus sexpustulatus
      • Brachinus oxygonus
      • Brachinus yunnanus
      • Brachinus vigilans
      • Brachinus abyssinicus
      • Brachinus sericeus
      • Brachinus gentilis
      • Brachinus chaudoirianus
      • Brachinus canaliculatus
      • Brachinus tigridis
      • Brachinus dieganus
      • Brachinus pallipes
      • Brachinus baeticus
      • A Carabid Beetle (Brachinus fumans)
      • Brachinus pelengensis
      • Bombardier Beetle (Brachinus cyanochroaticus)
      • Brachinus scutellatus
      • Brachinus xanthopleurus
      • Brachinus marginiventris
      • Brachinus hunanensis
      • Brachinus pateri
      • Brachinus deuvei
      • Brachinus lapidarius
      • Bombardier Beetle (Brachinus crepitans)
      • Brachinus bagdatensis
      • Brachinus nigripes
      • Brachinus knirschi
      • Brachinus orientalis
      • Brachinus chirriador
      • Brachinus enonensis
      • Brachinus immaculicornis
      • Brachinus mindanaoensis
      • Brachinus hexagrammus
      • Brachinus obliquetruncatus
      • A Carabid Beetle (Brachinus medius)
      • Brachinus schmidti
      • Brachinus nuristanus
      • Brachinus orestes
      • Brachinus bodemeyeri
      • Brachinus bipustulatus
      • Brachinus longulus
      • Brachinus fuscipennis
      • Brachinus stevensi
      • Brachinus primordialis
      • Brachinus ankarensis
      • Brachinus latipennis
      • Brachinus szetschuanensis
      • Brachinus flaviventris
      • Brachinus inops
      • Brachinus mexicanus
      • Brachinus lewisii
      • Brachinus stevensianus
      • Brachinus grandis
      • Brachinus puberulus
      • Brachinus incomptus
      • Brachinus intactus
      • Brachinus natalicus
      • Brachinus marleyi
      • Brachinus pecoudi
      • Brachinus votrubai
      • Brachinus explosus
      • Brachinus bolovenensis
      • Brachinus pallidipes
      • Brachinus charis
      • Brachinus pseudocruciatus
      • Brachinus solidipalpis
      • Brachinus lavaudeni
      • Brachinus thieleni
      • Brachinus scotomedes
      • Brachinus hoffmanni
      • Brachinus ciliatus
      • Brachinus vadoni
      • Brachinus flavicapillus
      • Brachinus ovalipennis
      • Brachinus foochowi
      • Brachinus ejaculans
      • Brachinus laevicostis
      • Brachinus kavanaughi
      • Brachinus pokharaensis
      • Brachinus seyrigi
      • Brachinus tetragrammus
      • Brachinus pectoralis
      • Brachinus ornatus
      • Brachinus johorensis
      • Brachinus consanguineus
      • Brachinus praestans
      • Brachinus barbarus
      • Brachinus tetracolon
      • Brachinus vicinus
      • Brachinus algoensis
      • Brachinus kalalovae
      • Brachinus vitticollis
      • Brachinus lewecki
      • Brachinus pallidus
      • Brachinus dawnaensis
      • Brachinus javalinopsis
      • Brachinus berytensis
      • Brachinus pygmaeus
      • Brachinus scotti
      • Brachinus congicus
      • Brachinus latus
      • Brachinus efflans
      • Brachinus flores
      • Brachinus angustatus
      • Brachinus suberbianus
      • Brachinus sichemita
      • Brachinus otini
      • Brachinus jucundus
      • Brachinus oneili
      • Brachinus explodens
      • Brachinus belyaevae
      • Brachinus apicalis
      • Brachinus olidus
      • Brachinus dilapsus
      • Brachinus formosanus
      • Brachinus forticostis
      • Brachinus variegatus
      • Brachinus vagus
      • Brachinus piceus
      • Brachinus inconditus
      • Brachinus nobilis
      • Brachinus subcostatus
      • Brachinus intermedius
      • Brachinus scitulus
      • Brachinus obliterus
      • A Ground Beetle (Brachinus janthinipennis)
      • Brachinus neglectus
      • Brachinus bellicosus
      • Brachinus modestus
      • Brachinus posticus
      • Brachinus sordidus
      • Brachinus philippinensis
      • Brachinus dissimilis
      • Brachinus annulicornis
      • Brachinus fuscicornis
      • Brachinus irakus
      • Brachinus aeger
      • Brachinus sulcipennis
      • Brachinus connectus
      • Brachinus suturatus
      • Brachinus grootaerti
      • Brachinus bigutticeps
      • Brachinus fasciatocollis
      • Brachinus exhalans
      • A Ground Beetle (Brachinus cordicollis)
      • Brachinus italicus
      • Brachinus quadriguttatus
    • Mastax → (52 species)
      • Mastax fulvonotata
      • Mastax nepalensis
      • Mastax philippina
      • Mastax pulchella
      • Mastax elegantula
      • Mastax klapperichi
      • Mastax congoensis
      • Mastax okavango
      • Mastax laeviceps
      • Mastax kivuensis
      • Mastax parreysii
      • Mastax extrema
      • Mastax alternans
      • Mastax albonotata
      • Mastax ornatella
      • Mastax ochraceonotata
      • Mastax ornata
      • Mastax rugiceps
      • Mastax thermarum
      • Mastax formosana
      • Mastax saganicola
      • Mastax raffrayi
      • Mastax poecila
      • Mastax carissima
      • Mastax latefasciata
      • Mastax striaticeps
      • Mastax histrio
      • Mastax subornatella
      • Mastax tratorius
      • Mastax tessmanni
      • Mastax senegalensis
      • Mastax euanthes
      • Mastax moesta
      • Mastax royi
      • Mastax burgeoni
      • Mastax annulata
      • Mastax humilis
      • Mastax louwerensi
      • Mastax florida
      • Mastax nana
      • Mastax gestroi
      • Mastax liebkei
      • Mastax vegeta
      • Mastax brittoni
      • Mastax sudanica
      • Mastax rawalpindi
      • Mastax hargreavesi
      • Mastax pygmaea
      • Mastax pakistana
      • Mastax fortesculpta
      • Mastax kulti
      • Mastax confusa
    • Styphlomerus → (47 species)
      • Styphlomerus ludicrus
      • Styphlomerus methneri
      • Styphlomerus lamottei
      • Styphlomerus titschacki
      • Styphlomerus exilis
      • Styphlomerus seyrigi
      • Styphlomerus postdilatatus
      • Styphlomerus montanus
      • Styphlomerus impressifrons
      • Styphlomerus plausibilis
      • Styphlomerus opacicollis
      • Styphlomerus puberulus
      • Styphlomerus korgei
      • Styphlomerus equestris
      • Styphlomerus paralleloides
      • Styphlomerus speciosus
      • Styphlomerus quadrimaculatus
      • Styphlomerus comoricus
      • Styphlomerus aulicus
      • Styphlomerus neavei
      • Styphlomerus gebieni
      • Styphlomerus ludricus
      • Styphlomerus fusciceps
      • Styphlomerus tellini
      • Styphlomerus cribricollis
      • Styphlomerus fuscifrons
      • Styphlomerus ovalipennis
      • Styphlomerus kamerunus
      • Styphlomerus sinus
      • Styphlomerus foveatus
      • Styphlomerus fallax
      • Styphlomerus kisantuus
      • Styphlomerus ciliatus
      • Styphlomerus timoriensis
      • Styphlomerus vittaticollis
      • Styphlomerus ruficeps
      • Styphlomerus flavus
      • Styphlomerus amplus
      • Styphlomerus piccolo
      • Styphlomerus fuscicollis
      • Styphlomerus undulatus
      • Styphlomerus kochi
      • Styphlomerus batesi
      • Styphlomerus burgeoni
      • Styphlomerus placidus
      • Styphlomerus renaudi
      • Styphlomerus sticticollis
    • Aptinus → (10 species)
      • Aptinus displosor
      • Aptinus alpinus
      • Aptinus pyrenaeus
      • Aptinus acutangulus
      • Aptinus lugubris
      • Aptinus cordicollis
      • Aptinus creticus
      • Aptinus bombarda
      • Aptinus hovorkai
      • Aptinus merditanus
    • Pheropsophus → (144 species)
      • Pheropsophus similis
      • Pheropsophus tristis
      • Pheropsophus nigerrimus
      • Pheropsophus minahassae
      • Pheropsophus platycephalus
      • Pheropsophus tamdaoensis
      • Pheropsophus androyanus
      • Pheropsophus planti
      • Pheropsophus congoensis
      • Pheropsophus beauvoisi
      • Pheropsophus iranicus
      • Pheropsophus nanodes
      • Pheropsophus andrewesi
      • Pheropsophus abbreviatus
      • Pheropsophus hispanicus
      • Pheropsophus sobrinus
      • Pheropsophus occipitalis
      • Pheropsophus langenhani
      • Pheropsophus amnicola
      • Pheropsophus kersteni
      • Pheropsophus maculicollis
      • Pheropsophus bimaculatus
      • Pheropsophus dux
      • Pheropsophus windjanae
      • Pheropsophus mashunus
      • Pheropsophus lafertei
      • Pheropsophus gregoryi
      • Pheropsophus straneoi
      • Pheropsophus nebulosus
      • Pheropsophus cinctus
      • Pheropsophus africanus
      • Pheropsophus exiguus
      • Pheropsophus bisulcatus
      • Pheropsophus brussinoi
      • Pheropsophus jessoensis
      • Pheropsophus yunnanensis
      • Pheropsophus picicollis
      • Pheropsophus aptinoides
      • Pheropsophus baehri
      • Pheropsophus impressicollis
      • Pheropsophus beckeri
      • Pheropsophus kalyakini
      • Pheropsophus uelensis
      • Pheropsophus malaisei
      • Pheropsophus rufimembris
      • Pheropsophus emarginatus
      • Pheropsophus ubomboensis
      • Pheropsophus livingstoni
      • Pheropsophus guanxiensis
      • Pheropsophus palmarum
      • Pheropsophus marginatus
      • Pheropsophus nyasae
      • Pheropsophus dregei
      • Pheropsophus abyssinicus
      • Pheropsophus gironieri
      • Pheropsophus lissoderus
      • Pheropsophus angolensis
      • Pheropsophus claudiae
      • Pheropsophus cardoni
      • Pheropsophus rolex
      • Pheropsophus tripustulatus
      • Pheropsophus biplagiatus
      • Pheropsophus laticostis
      • Pheropsophus nepalensis
      • Pheropsophus nigricollis
      • Pheropsophus siamensis
      • Pheropsophus obliquatus
      • Pheropsophus dilatatus
      • Pheropsophus aptinomorphus
      • Pheropsophus obliteratus
      • Pheropsophus jurinei
      • Pheropsophus borkuanus
      • Pheropsophus wolfi
      • Pheropsophus basiguttatus
      • Pheropsophus canis
      • Pheropsophus capensis
      • Pheropsophus halteri
      • Pheropsophus fastigiatus
      • Pheropsophus stenopterus
      • Pheropsophus bohemani
      • Pheropsophus chaudoiri
      • Pheropsophus catulus
      • Pheropsophus nigriventris
      • Pheropsophus consularis
      • Pheropsophus ecuadorensis
      • Pheropsophus bidoupensis
      • Pheropsophus javanus
      • Pheropsophus reductus
      • Pheropsophus rivierii
      • Pheropsophus alexandrae
      • Pheropsophus assimilis
      • Pheropsophus perroti
      • Pheropsophus riedeli
      • Pheropsophus acutecostatus
      • Pheropsophus fulminans
      • Pheropsophus prophylax
      • Pheropsophus humeralis
      • Pheropsophus aequinoctialis
      • Pheropsophus ngoclinhensis
      • Pheropsophus debauvii
      • Pheropsophus arcanus
      • Pheropsophus infantulus
      • Pheropsophus raffrayi
      • Pheropsophus delmastroi
      • Pheropsophus kolbei
      • Pheropsophus marginipennis
      • Pheropsophus pauliani
      • Pheropsophus verticalis
      • Pheropsophus baliothorax
      • Pheropsophus madagascariensis
      • Pheropsophus heathi
      • Pheropsophus pallidepunctatus
      • Pheropsophus discicollis
      • Pheropsophus transvaalensis
      • Pheropsophus parallelus
      • Pheropsophus cincticollis
      • Pheropsophus akhaensis
      • Pheropsophus globulicollis
      • Pheropsophus fumigatus
      • Pheropsophus bequaerti
      • Pheropsophus bipartitus
      • Pheropsophus chinensis
      • Pheropsophus katangensis
      • Pheropsophus krichna
      • Pheropsophus recticollis
      • Pheropsophus hassenteufeli
      • Pheropsophus catoirei
      • Pheropsophus scythropus
      • Pheropsophus hilaris
      • Pheropsophus senegalensis
      • Pheropsophus darwini
      • Pheropsophus marginicollis
      • Pheropsophus lineifrons
      • Pheropsophus uniformis
      • Pheropsophus capitatus
      • Pheropsophus balkei
      • Pheropsophus galloi
      • Pheropsophus gracilis
      • Pheropsophus longipennis
      • Pheropsophus dimidiatus
      • Pheropsophus pedes
      • Pheropsophus suensoni
      • Pheropsophus minor
      • Pheropsophus adrianae
    • Styphlodromus bicolor
  • Crepidogastrini → (110 species)
    • Crepidogaster holtzi
    • Crepidogaster dirrupta
    • Crepidogaster fortesculpta
    • Crepidogaster madecassa
    • Crepidogaster arnoldi
    • Crepidogaster specicola
    • Crepidogaster cicatricosa
    • Crepidogaster horni
    • Crepidogaster elongata
    • Crepidogaster didyma
    • Crepidogaster orophila
    • Crepidogaster aurasia
    • Crepidogaster insignis
    • Crepidogaster deuvei
    • Crepidogaster algoa
    • Crepidogaster subovicollis
    • Crepidogaster natalensis
    • Crepidogaster basilewskyi
    • Crepidogaster ferruginea
    • Crepidogaster jocquei
    • Crepidogaster mateui
    • Crepidogaster langenhani
    • Crepidogaster notulatipennis
    • Crepidogaster penrithae
    • Crepidogaster namaqua
    • Crepidogaster amieti
    • Crepidogaster louwerensi
    • Crepidogaster neglecta
    • Crepidogaster reducta
    • Crepidogaster garambae
    • Crepidogaster endroedyi
    • Crepidogaster infuscata
    • Crepidogaster insularis
    • Crepidogaster nonstriata
    • Crepidogaster elegantula
    • Crepidogaster kivuensis
    • Crepidogaster caeca
    • Crepidogaster dolini
    • Crepidogaster indica
    • Crepidogaster luberoensis
    • Crepidogaster phallica
    • Crepidogaster ceylanica
    • Crepidogaster iturica
    • Crepidogaster ruwenzorica
    • Crepidogaster aethiopica
    • Crepidogaster plagata
    • Crepidogaster marginicollis
    • Crepidogaster sambesiaca
    • Crepidogaster posticalis
    • Crepidogaster rufescens
    • Crepidogaster tendiculata
    • Crepidogaster longelineata
    • Crepidogaster oblata
    • Crepidogaster diluta
    • Crepidogaster kaboboensis
    • Crepidogaster protuberata
    • Crepidogaster malawica
    • Crepidogaster sansibarica
    • Crepidogaster kahuziana
    • Crepidogaster brincki
    • Crepidogaster venusta
    • Crepidogaster peyrierasi
    • Crepidogaster pauliani
    • Crepidogaster alticola
    • Crepidogaster angolana
    • Crepidogaster caffra
    • Crepidogaster muhavurae
    • Crepidogaster confusa
    • Crepidogaster monticola
    • Crepidogaster dissimilis
    • Crepidogaster costata
    • Crepidogaster delineata
    • Crepidogaster hessei
    • Crepidogaster bimaculata
    • Crepidogaster decolorata
    • Crepidogaster dollmani
    • Crepidogaster methneri
    • Crepidogaster ruandana
    • Crepidogaster ambreana
    • Crepidogaster transvaalensis
    • Crepidogaster damarensis
    • Crepidogaster leleupi
    • Crepidogaster laticollis
    • Crepidogaster kundelunguana
    • Crepidogaster franzi
    • Crepidogaster caffer
    • Crepidogaster lateralis
    • Crepidogaster kavanaughi
    • Crepidogaster mysorensis
    • Crepidogaster bambusicola
    • Crepidogaster oldeanica
    • Crepidogaster nyakagerana
    • Crepidogaster picipennis
    • Crepidogaster celsa
    • Crepidogaster humerata
    • Crepidogaster atrata
    • Crepidogaster humicola
    • Crepidogaster bioculata
    • Crepidogaster grossa
    • Crepidogaster occidentalis
    • Crepidogaster somalica
    • Crepidogaster ovicollis
    • Crepidogaster straneoi
    • Crepidogaster matonga
    • Crepidogaster funerula
    • Crepidogaster swazica
    • Crepidogaster schroederi
    • Crepidogaster natalica
    • Crepidogaster kochi
    • Crepidogaster portentosa

Other links

• Wikidata
• Wikipedia (language: en)
• Wikipedia (language: de)
• Wikipedia (language: es)
• Wikipedia (language: fr)
• Wikipedia (language: it)
• Wikipedia (language: pt)
• Wikipedia (language: ar)
• Wikipedia (language: nl)
• Wikipedia (language: fa)
• Wikipedia (language: tr)
• Encyclopedia of Life
• NCBI (genetics)"
• Open Tree of Life

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