Base of the evolutionary tree for Brachinini

← parent

• Brachinini (591 species)
  • Bombardier Beetle (Brachinus) → (337 species)
    • Brachinus hylaenus
    • Brachinus subnotatus
    • Brachinus elegans
    • Brachinus cinctellus
    • Brachinus albarracinus
    • Brachinus hamatus
    • Brachinus adustipennis
    • Brachinus tsara
    • Brachinus dilatipennis
    • Brachinus lombokensis
    • A Ground Beetle (Brachinus cyanipennis)
    • Brachinus tetraspilotus
    • A Carabid Beetle (Brachinus patruelis)
    • Brachinus melancholicus
    • Brachinus andreaei
    • Brachinus promontorii
    • A Ground Beetle (Brachinus alternans)
    • Brachinus costiger
    • Brachinus sexmaculatus
    • Brachinus afghanus
    • Brachinus sallei
    • Brachinus marginellus
    • Brachinus armiger
    • Brachinus peltastes
    • Brachinus tenuis
    • Brachinus atripensis
    • Brachinus eucosmus
    • Brachinus psophia
    • Brachinus illotus
    • Brachinus melanarthrus
    • Brachinus pachygaster
    • Brachinus macrocerus
    • Brachinus distans
    • Brachinus punctaticollis
    • Brachinus servillei
    • Brachinus limbellus
    • Brachinus dryas
    • Brachinus fageli
    • Brachinus hazardi
    • Brachinus sexnotatus
    • Brachinus circumtinctus
    • Brachinus americanus
    • Brachinus plagiatus
    • Brachinus aabaaba
    • Brachinus mesopotamicus
    • Brachinus rugipennis
    • A Ground Beetle (Brachinus tenuicollis)
    • Brachinus cruciatus
    • Brachinus connectoides
    • Brachinus oaxacensis
    • Brachinus limbicollis
    • Brachinus daliensis
    • Brachinus suensoni
    • Brachinus papua
    • Brachinus talyschensis
    • Brachinus testaceus
    • Brachinus xanthophryus
    • Brachinus brunneus
    • Brachinus cambodgensis
    • Brachinus cychroides
    • Brachinus midoli
    • Brachinus jakli
    • Brachinus micheli
    • Brachinus fulvipennis
    • Brachinus limbiger
    • A Ground Beetle (Brachinus fulminatus)
    • Brachinus favicollis
    • Brachinus repressus
    • Brachinus mauretanicus
    • Brachinus hirsutus
    • Brachinus dorsalis
    • Brachinus diffusus
    • Brachinus ghindanus
    • Brachinus flavus
    • Brachinus mactus
    • Brachinus chalchihuitlicue
    • Brachinus niger
    • Brachinus lateralis
    • Brachinus magyari
    • Brachinus variventris
    • Brachinus rikatlae
    • Brachinus evanescens
    • Brachinus humeralis
    • Brachinus obtusus
    • Brachinus ichabodopsis
    • Brachinus nigridorsis
    • Brachinus minor
    • Brachinus gebhardis
    • Brachinus methneri
    • Brachinus cibolensis
    • Brachinus sicardi
    • Brachinus lesnei
    • Brachinus kollari
    • Brachinus dilottii
    • Brachinus microrrhabdus
    • Brachinus bantimurungensis
    • Brachinus maublanci
    • Brachinus imporcitis
    • Brachinus alexandri
    • Brachinus suturellus
    • Brachinus collarti
    • Brachinus atramentarius
    • Streaked Bombardier (Brachinus sclopeta)
    • Brachinus madecassus
    • Brachinus langenhani
    • Brachinus fukienensis
    • Brachinus nigricans
    • Brachinus sonorous
    • Brachinus luzonicus
    • Brachinus frontalis
    • Brachinus nigrovirescens
    • Brachinus quadripennis
    • Brachinus puncticollis
    • Brachinus boeticus
    • Brachinus tianshanicus
    • Brachinus costulipennis
    • Brachinus newberryi
    • Brachinus sublaevis
    • Brachinus turnai
    • Brachinus proximus
    • Brachinus leprieuri
    • Brachinus alexiguus
    • Brachinus lethierryi
    • Brachinus palawanensis
    • Brachinus aeneicostis
    • Brachinus demoulini
    • Brachinus flavipes
    • Brachinus abbreviatus
    • Brachinus constrictus
    • Brachinus perplexus
    • Brachinus vulcanoides
    • Brachinus bilineatus
    • Brachinus reyi
    • Brachinus brittoni
    • Brachinus pictus
    • Brachinus turkestanicus
    • Brachinus hubeiensis
    • Brachinus mobilis
    • Brachinus microamericanus
    • Brachinus agraphus
    • Brachinus bayardi
    • Brachinus dilatatus
    • Brachinus velutinus
    • Brachinus azureipennis
    • Brachinus nigricornis
    • Brachinus truncatulus
    • Brachinus elegantulus
    • Brachinus phaeocerus
    • Brachinus tetrastigma
    • Brachinus imperialensis
    • Brachinus scriptus
    • Brachinus laetus
    • Brachinus kansanus
    • Brachinus brevicollis
    • Brachinus cinctipennis
    • Brachinus costatulus
    • Brachinus simulans
    • Brachinus stenoderus
    • Brachinus costipennis
    • Brachinus merkli
    • Brachinus arboreus
    • Brachinus immarginatus
    • Brachinus bruchi
    • Brachinus dalatensis
    • Brachinus solidus
    • Brachinus stappersi
    • Brachinus geniculatus
    • Brachinus batuensis
    • Brachinus caffer
    • Brachinus chinensis
    • Brachinus paviei
    • Brachinus suturalis
    • Brachinus viridipennis
    • Brachinus elongatulus
    • Brachinus exquisitus
    • Brachinus texanus
    • Brachinus stygius
    • Brachinus perrieri
    • Brachinus conformis
    • Brachinus longipalpis
    • A Carabid Beetle (Brachinus ovipennis)
    • Brachinus olgae
    • Brachinus andalusiacus
    • Brachinus galactoderus
    • Brachinus capnicus
    • Brachinus catalonicus
    • Brachinus oblongus
    • Brachinus genicularis
    • Brachinus somereni
    • Brachinus aristokrates
    • Brachinus sexpustulatus
    • Brachinus oxygonus
    • Brachinus yunnanus
    • Brachinus vigilans
    • Brachinus abyssinicus
    • Brachinus sericeus
    • Brachinus gentilis
    • Brachinus chaudoirianus
    • Brachinus canaliculatus
    • Brachinus tigridis
    • Brachinus dieganus
    • Brachinus pallipes
    • Brachinus baeticus
    • A Carabid Beetle (Brachinus fumans)
    • Brachinus pelengensis
    • Bombardier Beetle (Brachinus cyanochroaticus)
    • Brachinus scutellatus
    • Brachinus xanthopleurus
    • Brachinus marginiventris
    • Brachinus hunanensis
    • Brachinus pateri
    • Brachinus deuvei
    • Brachinus lapidarius
    • Bombardier Beetle (Brachinus crepitans)
    • Brachinus bagdatensis
    • Brachinus nigripes
    • Brachinus knirschi
    • Brachinus orientalis
    • Brachinus chirriador
    • Brachinus enonensis
    • Brachinus immaculicornis
    • Brachinus mindanaoensis
    • Brachinus hexagrammus
    • Brachinus obliquetruncatus
    • A Carabid Beetle (Brachinus medius)
    • Brachinus schmidti
    • Brachinus nuristanus
    • Brachinus orestes
    • Brachinus bodemeyeri
    • Brachinus bipustulatus
    • Brachinus longulus
    • Brachinus fuscipennis
    • Brachinus stevensi
    • Brachinus primordialis
    • Brachinus ankarensis
    • Brachinus latipennis
    • Brachinus szetschuanensis
    • Brachinus flaviventris
    • Brachinus inops
    • Brachinus mexicanus
    • Brachinus lewisii
    • Brachinus stevensianus
    • Brachinus grandis
    • Brachinus puberulus
    • Brachinus incomptus
    • Brachinus intactus
    • Brachinus natalicus
    • Brachinus marleyi
    • Brachinus pecoudi
    • Brachinus votrubai
    • Brachinus explosus
    • Brachinus bolovenensis
    • Brachinus pallidipes
    • Brachinus charis
    • Brachinus pseudocruciatus
    • Brachinus solidipalpis
    • Brachinus lavaudeni
    • Brachinus thieleni
    • Brachinus scotomedes
    • Brachinus hoffmanni
    • Brachinus ciliatus
    • Brachinus vadoni
    • Brachinus flavicapillus
    • Brachinus ovalipennis
    • Brachinus foochowi
    • Brachinus ejaculans
    • Brachinus laevicostis
    • Brachinus kavanaughi
    • Brachinus pokharaensis
    • Brachinus seyrigi
    • Brachinus tetragrammus
    • Brachinus pectoralis
    • Brachinus ornatus
    • Brachinus johorensis
    • Brachinus consanguineus
    • Brachinus praestans
    • Brachinus barbarus
    • Brachinus tetracolon
    • Brachinus vicinus
    • Brachinus algoensis
    • Brachinus kalalovae
    • Brachinus vitticollis
    • Brachinus lewecki
    • Brachinus pallidus
    • Brachinus dawnaensis
    • Brachinus javalinopsis
    • Brachinus berytensis
    • Brachinus pygmaeus
    • Brachinus scotti
    • Brachinus congicus
    • Brachinus latus
    • Brachinus efflans
    • Brachinus flores
    • Brachinus angustatus
    • Brachinus suberbianus
    • Brachinus sichemita
    • Brachinus otini
    • Brachinus jucundus
    • Brachinus oneili
    • Brachinus explodens
    • Brachinus belyaevae
    • Brachinus apicalis
    • Brachinus olidus
    • Brachinus dilapsus
    • Brachinus formosanus
    • Brachinus forticostis
    • Brachinus variegatus
    • Brachinus vagus
    • Brachinus piceus
    • Brachinus inconditus
    • Brachinus nobilis
    • Brachinus subcostatus
    • Brachinus intermedius
    • Brachinus scitulus
    • Brachinus obliterus
    • A Ground Beetle (Brachinus janthinipennis)
    • Brachinus neglectus
    • Brachinus bellicosus
    • Brachinus modestus
    • Brachinus posticus
    • Brachinus sordidus
    • Brachinus philippinensis
    • Brachinus dissimilis
    • Brachinus annulicornis
    • Brachinus fuscicornis
    • Brachinus irakus
    • Brachinus aeger
    • Brachinus sulcipennis
    • Brachinus connectus
    • Brachinus suturatus
    • Brachinus grootaerti
    • Brachinus bigutticeps
    • Brachinus fasciatocollis
    • Brachinus exhalans
    • A Ground Beetle (Brachinus cordicollis)
    • Brachinus italicus
    • Brachinus quadriguttatus
  • Mastax → (52 species)
    • Mastax fulvonotata
    • Mastax nepalensis
    • Mastax philippina
    • Mastax pulchella
    • Mastax elegantula
    • Mastax klapperichi
    • Mastax congoensis
    • Mastax okavango
    • Mastax laeviceps
    • Mastax kivuensis
    • Mastax parreysii
    • Mastax extrema
    • Mastax alternans
    • Mastax albonotata
    • Mastax ornatella
    • Mastax ochraceonotata
    • Mastax ornata
    • Mastax rugiceps
    • Mastax thermarum
    • Mastax formosana
    • Mastax saganicola
    • Mastax raffrayi
    • Mastax poecila
    • Mastax carissima
    • Mastax latefasciata
    • Mastax striaticeps
    • Mastax histrio
    • Mastax subornatella
    • Mastax tratorius
    • Mastax tessmanni
    • Mastax senegalensis
    • Mastax euanthes
    • Mastax moesta
    • Mastax royi
    • Mastax burgeoni
    • Mastax annulata
    • Mastax humilis
    • Mastax louwerensi
    • Mastax florida
    • Mastax nana
    • Mastax gestroi
    • Mastax liebkei
    • Mastax vegeta
    • Mastax brittoni
    • Mastax sudanica
    • Mastax rawalpindi
    • Mastax hargreavesi
    • Mastax pygmaea
    • Mastax pakistana
    • Mastax fortesculpta
    • Mastax kulti
    • Mastax confusa
  • Styphlomerus → (47 species)
    • Styphlomerus ludicrus
    • Styphlomerus methneri
    • Styphlomerus lamottei
    • Styphlomerus titschacki
    • Styphlomerus exilis
    • Styphlomerus seyrigi
    • Styphlomerus postdilatatus
    • Styphlomerus montanus
    • Styphlomerus impressifrons
    • Styphlomerus plausibilis
    • Styphlomerus opacicollis
    • Styphlomerus puberulus
    • Styphlomerus korgei
    • Styphlomerus equestris
    • Styphlomerus paralleloides
    • Styphlomerus speciosus
    • Styphlomerus quadrimaculatus
    • Styphlomerus comoricus
    • Styphlomerus aulicus
    • Styphlomerus neavei
    • Styphlomerus gebieni
    • Styphlomerus ludricus
    • Styphlomerus fusciceps
    • Styphlomerus tellini
    • Styphlomerus cribricollis
    • Styphlomerus fuscifrons
    • Styphlomerus ovalipennis
    • Styphlomerus kamerunus
    • Styphlomerus sinus
    • Styphlomerus foveatus
    • Styphlomerus fallax
    • Styphlomerus kisantuus
    • Styphlomerus ciliatus
    • Styphlomerus timoriensis
    • Styphlomerus vittaticollis
    • Styphlomerus ruficeps
    • Styphlomerus flavus
    • Styphlomerus amplus
    • Styphlomerus piccolo
    • Styphlomerus fuscicollis
    • Styphlomerus undulatus
    • Styphlomerus kochi
    • Styphlomerus batesi
    • Styphlomerus burgeoni
    • Styphlomerus placidus
    • Styphlomerus renaudi
    • Styphlomerus sticticollis
  • Aptinus → (10 species)
    • Aptinus displosor
    • Aptinus alpinus
    • Aptinus pyrenaeus
    • Aptinus acutangulus
    • Aptinus lugubris
    • Aptinus cordicollis
    • Aptinus creticus
    • Aptinus bombarda
    • Aptinus hovorkai
    • Aptinus merditanus
  • Pheropsophus → (144 species)
    • Pheropsophus similis
    • Pheropsophus tristis
    • Pheropsophus nigerrimus
    • Pheropsophus minahassae
    • Pheropsophus platycephalus
    • Pheropsophus tamdaoensis
    • Pheropsophus androyanus
    • Pheropsophus planti
    • Pheropsophus congoensis
    • Pheropsophus beauvoisi
    • Pheropsophus iranicus
    • Pheropsophus nanodes
    • Pheropsophus andrewesi
    • Pheropsophus abbreviatus
    • Pheropsophus hispanicus
    • Pheropsophus sobrinus
    • Pheropsophus occipitalis
    • Pheropsophus langenhani
    • Pheropsophus amnicola
    • Pheropsophus kersteni
    • Pheropsophus maculicollis
    • Pheropsophus bimaculatus
    • Pheropsophus dux
    • Pheropsophus windjanae
    • Pheropsophus mashunus
    • Pheropsophus lafertei
    • Pheropsophus gregoryi
    • Pheropsophus straneoi
    • Pheropsophus nebulosus
    • Pheropsophus cinctus
    • Pheropsophus africanus
    • Pheropsophus exiguus
    • Pheropsophus bisulcatus
    • Pheropsophus brussinoi
    • Pheropsophus jessoensis
    • Pheropsophus yunnanensis
    • Pheropsophus picicollis
    • Pheropsophus aptinoides
    • Pheropsophus baehri
    • Pheropsophus impressicollis
    • Pheropsophus beckeri
    • Pheropsophus kalyakini
    • Pheropsophus uelensis
    • Pheropsophus malaisei
    • Pheropsophus rufimembris
    • Pheropsophus emarginatus
    • Pheropsophus ubomboensis
    • Pheropsophus livingstoni
    • Pheropsophus guanxiensis
    • Pheropsophus palmarum
    • Pheropsophus marginatus
    • Pheropsophus nyasae
    • Pheropsophus dregei
    • Pheropsophus abyssinicus
    • Pheropsophus gironieri
    • Pheropsophus lissoderus
    • Pheropsophus angolensis
    • Pheropsophus claudiae
    • Pheropsophus cardoni
    • Pheropsophus rolex
    • Pheropsophus tripustulatus
    • Pheropsophus biplagiatus
    • Pheropsophus laticostis
    • Pheropsophus nepalensis
    • Pheropsophus nigricollis
    • Pheropsophus siamensis
    • Pheropsophus obliquatus
    • Pheropsophus dilatatus
    • Pheropsophus aptinomorphus
    • Pheropsophus obliteratus
    • Pheropsophus jurinei
    • Pheropsophus borkuanus
    • Pheropsophus wolfi
    • Pheropsophus basiguttatus
    • Pheropsophus canis
    • Pheropsophus capensis
    • Pheropsophus halteri
    • Pheropsophus fastigiatus
    • Pheropsophus stenopterus
    • Pheropsophus bohemani
    • Pheropsophus chaudoiri
    • Pheropsophus catulus
    • Pheropsophus nigriventris
    • Pheropsophus consularis
    • Pheropsophus ecuadorensis
    • Pheropsophus bidoupensis
    • Pheropsophus javanus
    • Pheropsophus reductus
    • Pheropsophus rivierii
    • Pheropsophus alexandrae
    • Pheropsophus assimilis
    • Pheropsophus perroti
    • Pheropsophus riedeli
    • Pheropsophus acutecostatus
    • Pheropsophus fulminans
    • Pheropsophus prophylax
    • Pheropsophus humeralis
    • Pheropsophus aequinoctialis
    • Pheropsophus ngoclinhensis
    • Pheropsophus debauvii
    • Pheropsophus arcanus
    • Pheropsophus infantulus
    • Pheropsophus raffrayi
    • Pheropsophus delmastroi
    • Pheropsophus kolbei
    • Pheropsophus marginipennis
    • Pheropsophus pauliani
    • Pheropsophus verticalis
    • Pheropsophus baliothorax
    • Pheropsophus madagascariensis
    • Pheropsophus heathi
    • Pheropsophus pallidepunctatus
    • Pheropsophus discicollis
    • Pheropsophus transvaalensis
    • Pheropsophus parallelus
    • Pheropsophus cincticollis
    • Pheropsophus akhaensis
    • Pheropsophus globulicollis
    • Pheropsophus fumigatus
    • Pheropsophus bequaerti
    • Pheropsophus bipartitus
    • Pheropsophus chinensis
    • Pheropsophus katangensis
    • Pheropsophus krichna
    • Pheropsophus recticollis
    • Pheropsophus hassenteufeli
    • Pheropsophus catoirei
    • Pheropsophus scythropus
    • Pheropsophus hilaris
    • Pheropsophus senegalensis
    • Pheropsophus darwini
    • Pheropsophus marginicollis
    • Pheropsophus lineifrons
    • Pheropsophus uniformis
    • Pheropsophus capitatus
    • Pheropsophus balkei
    • Pheropsophus galloi
    • Pheropsophus gracilis
    • Pheropsophus longipennis
    • Pheropsophus dimidiatus
    • Pheropsophus pedes
    • Pheropsophus suensoni
    • Pheropsophus minor
    • Pheropsophus adrianae
  • Styphlodromus bicolor

Other links

• Wikidata
• Wikipedia (language: es)
• Wikipedia (language: pt)
• Encyclopedia of Life
• NCBI (genetics)"
• Open Tree of Life

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