Base of the evolutionary tree for Pupilloidea

← parent

• Pupilloidea (1340 species)
  • Gastrocoptidae → (212 species)
    • Gastrocopta → (151 species)
      • Sinalbinula pleimesi
      • Sinalbinula recondida
      • Sinalbinula orientalis
      • Sinalbinula armigerella
      • Sinalbinula ovatula
      • Sinalbinula kamberae
      • Sinalbinula theeli
      • Sinalbinula microsoma
      • Sinalbinula niobe
      • Sinalbinula meridionalis
      • Gastrocopta disjuncta
      • Gastrocopta reibischi
      • Gastrocopta recondita
      • Gastrocopta polyptyx
      • Gastrocopta calumniosa
      • Gastrocopta gemina
      • Gastrocopta theeli
      • Gastrocopta polonica
      • Gastrocopta biruleica
      • Gastrocopta proarmifera
      • Gastrocopta chichijimana
      • Gastrocopta duncana
      • Gastrocopta moravica
      • Apache snaggletooth (Gastrocopta cochisensis)
      • sluice snaggletooth (Gastrocopta ashmuni)
      • Gastrocopta boninensis
      • Gastrocopta damarica
      • Sonoran snaggletooth (Gastrocopta prototypus)
      • Gulf Coast snaggletooth (Gastrocopta riparia)
      • Gastrocopta ejecta
      • plains snaggletooth (Gastrocopta abbreviata)
      • Gastrocopta armigerella
      • Gastrocopta wolfii
      • Gastrocopta subumbilicata
      • Gastrocopta aliciae
      • Gastrocopta rixfordi
      • Gastrocopta kintlana
      • Gastrocopta klunzingeri
      • Gastrocopta flocculus
      • Gastrocopta falcis
      • Gastrocopta ooi
      • tapered snaggletooth (Gastrocopta rupicola)
      • Gastrocopta hedleyi
      • Gastrocopta patagonica
      • Gastrocopta akokala
      • Gastrocopta bathyodon
      • Gastrocopta avanica
      • Gastrocopta semiclausa
      • Gastrocopta floridana
      • Gastrocopta microbus
      • Gastrocopta anterides
      • Gastrocopta herrerai
      • Gastrocopta oviforma
      • Gastrocopta microscopica
      • Gastrocopta magna
      • Gastrocopta tavennerensis
      • slim snaggletooth (Gastrocopta pellucida)
      • bluegrass snaggletooth (Gastrocopta clappi)
      • comb snaggletooth (Gastrocopta pentodon)
      • Gastrocopta sharae
      • Gastrocopta itaboraiensis
      • Gastrocopta lamellidens
      • Gastrocopta primitiva
      • Gastrocopta obesa
      • Gastrocopta curyomphata
      • Gastrocopta rhenana
      • Gastrocopta bannertonensis
      • Gastrocopta sterkiana
      • Gastrocopta hummelincki
      • Gastrocopta quadriplicata
      • bark snaggletooth (Gastrocopta corticaria)
      • Gastrocopta iheringi
      • Gastrocopta thomasseti
      • Gastrocopta jasamanovi
      • Gastrocopta christineae
      • Gastrocopta acarus
      • Rio Grande snaggletooth (Gastrocopta riograndensis)
      • Gastrocopta amoena
      • wandering snag (Gastrocopta servilis)
      • Gastrocopta kedeica
      • Gastrocopta rogersensis
      • Gastrocopta oblonga
      • Gastrocopta margaretae
      • Gastrocopta leonardi
      • Gastrocopta russelli
      • Gastrocopta barbadensis
      • Gastrocopta antinorii
      • Gastrocopta clausa
      • Gastrocopta ukrainica
      • white snaggletooth (Gastrocopta tappaniana)
      • cross snaggletooth (Gastrocopta quadridens)
      • Gastrocopta conica
      • Gastrocopta montana
      • Gastrocopta franki
      • Gastrocopta skiphica
      • Erie snaggletooth (Gastrocopta carnegiei)
      • Gastrocopta dahli
      • Gastrocopta sagittaria
      • Gastrocopta kyrminiensis
      • Gastrocopta palmira
      • Gastrocopta larapinta
      • Gastrocopta pediculus
      • Gastrocopta krestnikovi
      • Gastrocopta coreana
      • Gastrocopta paracristata
      • bottleneck snaggletooth (Gastrocopta contracta)
      • Gastrocopta obstructa
      • armed snaggletooth (Gastrocopta armifera)
      • Gastrocopta pontifex
      • Gastrocopta coniculus
      • Gastrocopta intorta
      • Gastrocopta farcimen
      • Gastrocopta steklovi
      • Gastrocopta seignaciana
      • wing snaggletooth (Gastrocopta procera)
      • lambda snaggletooth (Gastrocopta holzingeri)
      • montane snaggletooth (Gastrocopta pilsbryana)
      • Gastrocopta huttoniana
      • Gastrocopta mimula
      • shortneck snaggletooth (Gastrocopta dalliana)
      • Great Lakes snaggletooth (Gastrocopta similis)
      • Gastrocopta bugensis
      • Gastrocopta cordillerae
      • Gastrocopta stupefaciens
      • Gastrocopta aequatorialis
      • Gastrocopta minuscula
      • Gastrocopta munita
      • Gastrocopta mezzalirai
      • Gastrocopta strangeana
      • Gastrocopta jeanneli
      • Gastrocopta perfida
      • Gastrocopta praebaicalensis
      • Gastrocopta rexroadensis
      • Gastrocopta curacoana
      • Gastrocopta lienardiana
      • Gastrocopta zamankulense
      • Gastrocopta mussoni
      • Gastrocopta macdonnelli
      • crested snaggletooth (Gastrocopta cristata)
      • Gastrocopta haessleini
      • Gastrocopta subzamankulense
      • Gastrocopta kamberae
      • Gastrocopta geminidens
      • Gastrocopta eudeli
      • Gastrocopta abyssifluminis
      • Gastrocopta rybakovi
      • Gastrocopta allyni
      • Gastrocopta euryomphala
      • Ruidoso snaggletooth (Gastrocopta ruidosensis)
      • Gastrocopta kazachica
      • Gastrocopta gularis
    • Tonkinospira → (5 species)
      • Tonkinospira tomasini
      • Tonkinospira depressa
      • Tonkinospira pauperrima
      • Tonkinospira defixa
      • Tonkinospira pulverea
    • Chaenaxis → (2 species)
      • hollow tuba (Chaenaxis tuba)
      • Chaenaxis sonorensis
    • Pseudostreptaxis → (2 species)
      • Pseudostreptaxis azpeitiae
      • Pseudostreptaxis harli
    • Boysidia → (24 species)
      • Boysidia novemdentata
      • Boysidia dorsata
      • Boysidia gracilis
      • Boysidia hunana
      • Boysidia jinpingensis
      • Boysidia paini
      • Boysidia tamtouriana
      • Boysidia strophostoma
      • Boysidia robusta
      • Boysidia chiangmaiensis
      • Boysidia ringens
      • Boysidia kelantanense
      • Boysidia orientalis
      • Boysidia paviei
      • Boysidia phatangensis
      • Boysidia procera
      • Boysidia tholus
      • Boysidia megaphona
      • Boysidia elephas
      • Boysidia perigyra
      • Boysidia pahpetensis
      • Boysidia lamothei
      • Boysidia salpinx
      • Boysidia terae
    • Angustopila → (11 species)
      • Angustopila tamlod
      • Angustopila singuladentis
      • Angustopila szekeresi
      • Angustopila stochi
      • Angustopila fabella
      • Angustopila concava
      • Angustopila neglecta
      • Angustopila dominikae
      • Angustopila elevata
      • Angustopila huoyani
      • Angustopila subelevata
    • Bensonella → (2 species)
      • Bensonella karoensis
      • Bensonella plicidens
    • Acinolaemus → (7 species)
      • Acinolaemus rectus
      • Acinolaemus carcharodon
      • Acinolaemus colpodon
      • Acinolaemus sphinctinion
      • Acinolaemus ptychochilus
      • Acinolaemus stenopus
      • Acinolaemus rhamphodon
    • Montapiculus → (2 species)
      • Montapiculus proboscidea
      • Montapiculus pyramidalis
    • Antroapiculus pendulus
    • Gibbulina venusta
    • Ptychalaea dedecora
    • Boysia boysii
    • Campolaemus perexilis
    • Dentisphaera maxema
  • Lauriidae → (79 species)
    • Leiostyla → (60 species)
      • Leiostyla recta
      • Leiostyla lamellosa
      • Leiostyla eikenboomi
      • Leiostyla superba
      • Leiostyla iranica
      • Leiostyla silicea
      • Leiostyla schweigeri
      • Leiostyla espigaoensis
      • Leiostyla crassilabris
      • Leiostyla loweana
      • Leiostyla heterodon
      • Leiostyla minax
      • Leiostyla subcorneocostata
      • Leiostyla pontica
      • Leiostyla rugulosa
      • Leiostyla paphlagonica
      • Leiostyla beatae
      • Leiostyla honesta
      • Leiostyla falknerorum
      • Leiostyla paphalagonica
      • Leiostyla castanea
      • Leiostyla anglica
      • Leiostyla vermiculosa
      • Leiostyla tesselata
      • Leiostyla fuscidula
      • Leiostyla millegrana
      • Leiostyla colvillei
      • Leiostyla mica
      • Leiostyla caucasica
      • Leiostyla wollastoni
      • Leiostyla calathiscus
      • Leiostyla concinna
      • Leiostyla laurinea
      • Leiostyla subcornecostata
      • Leiostyla vitrea
      • Leiostyla cassidula
      • Leiostyla degenerata
      • Leiostyla taeniata
      • Madeiran land snail (Leiostyla corneocostata)
      • Leiostyla monticola
      • Leiostyla capellinii
      • Leiostyla piserai
      • Leiostyla fusca
      • Leiostyla relevata
      • Leiostyla filicum
      • Leiostyla nemethi
      • Leiostyla cheilogona
      • Leiostyla abbreviata
      • Leiostyla ferraria
      • Leiostyla vincta
      • Leiostyla rectidentata
      • Leiostyla albina
      • Leiostyla irrigua
      • Leiostyla gibba
      • Leiostyla simulator
      • Leiostyla zonifera
      • Leiostyla sphinctostoma
      • Leiostyla arborea
      • Leiostyla priscilla
      • Leiostyla cassida
    • Lauria → (18 species)
      • Lauria reischuetzi
      • Chrysalis Snail (Lauria cylindracea)
      • Lauria longa
      • Lauria desiderata
      • Lauria tingitana
      • Lauria bourbonensis
      • Lauria cryptoplax
      • Lauria wouramboulchiensis
      • Lauria sempronii
      • Lauria dadion
      • Lauria leprevieri
      • Lauria farquhari
      • Lauria fanalensis
      • Lauria gomerensis
      • Lauria bequaerti
      • Lauria umbilica
      • Lauria karisimbiensis
      • Lauria fasciolata
    • Hemilauria limnaeana
  • grass snails (Valloniidae) → (58 species)
    • Vallonia → (32 species)
      • Vallonia kootenayorum
      • Vallonia berryi
      • Vallonia patens
      • Vallonia girauxae
      • multirib vallonia (Vallonia gracilicosta)
      • ribbed grass snail (Vallonia costata)
      • Vallonia enniensis
      • indecisive vallonia (Vallonia albula)
      • Vallonia peteri
      • Vallonia hipparionum
      • Vallonia ladacensis
      • Vallonia brochoni
      • Vallonia sandbergeri
      • Vallonia asiatica
      • Vallonia tenuilabris
      • Vallonia pallaryi
      • Terre-Neuve vallonia (Vallonia terraenovae)
      • Vallonia suevica
      • trumpet vallonia (Vallonia parvula)
      • Vallonia zaru
      • Vallonia sparnacensis
      • eccentric grass snail (Vallonia excentrica)
      • Vallonia chinensis
      • Vallonia mionecton
      • smooth grass snail (Vallonia pulchella)
      • Vallonia major
      • Vallonia ranovi
      • Vallonia declivis
      • Vallonia pulchellula
      • silky vallonia (Vallonia cyclophorella)
      • thin-lip vallonia (Vallonia perspectiva)
      • Vallonia kamtschatica
    • Acanthinula → (15 species)
      • Acanthinula expatriata
      • Acanthinula sublabyrinthica
      • Acanthinula nanxiongensis
      • Acanthinula paronae
      • Acanthinula azorica
      • Acanthinula stampinensis
      • Acanthinula cenchridium
      • Acanthinula straeleni
      • Acanthinula stueri
      • Acanthinula armoricensis
      • Acanthinula spinifera
      • Acanthinula lamellata
      • Acanthinula dumasi
      • Acanthinula aculeata
      • Acanthinula archiaci
    • Plagyrona → (2 species)
      • Plagyrona placida
      • Plagyrona angusta
    • Nanxiongospira → (3 species)
      • Nanxiongospira uniptychia
      • Nanxiongospira nobilis
      • Nanxiongospira meiguanensis
    • Spermodea → (2 species)
      • Spermodea monas
      • Plaited snail (Spermodea lamellata)
    • Shanghuspira costata
    • boreal top (Zoogenetes harpa)
    • Gittenbergia sororcula
    • Sexlamellospira austriaca
  • snaggletooth snails (Chondrinidae) → (89 species)
    • Chondrina → (47 species)
      • Chondrina tatrica
      • Chondrina kobelti
      • Chondrina arcadica
      • Chondrina multidentata
      • Chondrina aguilari
      • Chondrina dertosensis
      • Chondrina caligunosus
      • Chondrina massotiana
      • Chondrina keltiensis
      • Chondrina spelta
      • Chondrina megacheilos
      • Chondrina arigonis
      • Chondrina ascendens
      • Chondrina farinesii
      • Chondrina soleri
      • Chondrina centralis
      • Chondrina granatensis
      • Chondrina michaudi
      • Chondrina feneriensis
      • Chondrina jumillensis
      • Chondrina klemmi
      • Chondrina bigorriensis
      • Chondrina bergomensis
      • Chondrina avenacea
      • Chondrina kobeltoides
      • Chondrina guiraoensis
      • Chondrina marjae
      • Chondrina lusitanica
      • Chondrina pseudoteresae
      • Chondrina gasulli
      • Chondrina gavirai
      • Chondrina gerhardi
      • Chondrina arigonoides
      • Chondrina muehlfeldti
      • Chondrina gomezi
      • Chondrina maginensis
      • Chondrina altimirai
      • Chondrina falkneri
      • Chondrina calpica
      • Chondrina generosensis
      • Chondrina marmouchana
      • Chondrina ingae
      • Chondrina tenuimarginata
      • Chondrina amphorula
      • Chondrina oligodonta
      • Chondrina ripkeni
      • Chondrina pseudavenacea
    • Granopupa → (5 species)
      • Granopupa somalensis
      • Granopupa persica
      • Granopupa klemmi
      • Granopupa hagenmulleri
      • Granopupa granum
    • Granaria → (8 species)
      • Granaria moedlingensis
      • Granaria frumentum
      • Granaria lapidaria
      • Granaria stabili
      • Granaria fustis
      • Granaria variabilis
      • Granaria arabica
      • Granaria persica
    • Rupestrella → (7 species)
      • Rupestrella dupotetii
      • Rupestrella homala
      • Rupestrella jaeckeli
      • Rupestrella rhodia
      • Rupestrella occulta
      • Rupestrella philippii
      • Rupestrella rupestris
    • Abida → (15 species)
      • Abida attenuata
      • Abida bigerrensis
      • Abida occidentalis
      • Abida polyodon
      • Abida cadica
      • Abida gittenbergeri
      • Abida partioti
      • Abida vasconica
      • Abida secale
      • Abida pyrenaearia
      • Abida polydon
      • Abida oryza
      • Abida cylindrica
      • Abida servasensis
      • Abida vouastensis
    • Solatopupa → (6 species)
      • Solatopupa similis
      • Solatopupa pallida
      • Solatopupa juliana
      • Solatopupa cianensis
      • Solatopupa psarolena
      • Solatopupa guidoni
    • Granaria braunii
  • Strobilopsidae → (68 species)
    • Strobilops → (37 species)
      • Discostrobilops incerta
      • Discostrobilops pericarinata
      • Discostrobilops robusta
      • Discostrobilops diploptycha
      • Strobila labyrinthica
      • Strobilops sparsicostatus
      • Strobilops steklovi
      • Strobilops monilia
      • eightfold pinecone (Strobilops affinis)
      • Strobilops menardi
      • Strobilops ukrainicaeformis
      • Strobilops stenostoma
      • Strobilops brasiliana
      • southern pinecone (Strobilops texasianus)
      • Strobilops veracruzensis
      • flattened pinecone (Strobilops hubbardi)
      • Strobilops helleri
      • Strobilops pseudolabyrinthicus
      • Strobilops salvini
      • bronze pinecone (Strobilops aeneus)
      • Strobilops desmarestinus
      • Strobilops lonsdalei
      • Strobilops piratica
      • Strobilops hannai
      • Strobilops morsei
      • Strobilops luosigangensis
      • Strobilops multidenticulata
      • Strobilops menneri
      • Strobilops strebeli
      • Strobilops labryinthica
      • Strobilops pilsbryi
      • Strobilops labyrinthiculus
      • Strobilops labyrinthicus
      • Strobilops headonensis
      • Strobilops californica
      • Strobilops lata
      • Strobilops sinaloa
    • Enteroplax → (8 species)
      • Enteroplax quadrasi
      • Enteroplax misoolensis
      • Enteroplax polyptychia
      • Enteroplax luosigangensis
      • Enteroplax parvus
      • Enteroplax stenostoma
      • Enteroplax trochospira
      • Enteroplax dumogensis
    • Prionolabium → (2 species)
      • Prionolabium decilamellatum
      • Prionolabium polyptychium
    • Eostrobilops → (17 species)
      • Eostrobilops diodontina
      • Eostrobilops zijinshanicus
      • Eostrobilops kongoensis
      • Eostrobilops subconoideus
      • Eostrobilops caucasica
      • Eostrobilops sinensis
      • Eostrobilops nipponica
      • Eostrobilops hirasei
      • Eostrobilops yaeyamensis
      • Eostrobilops dollfusi
      • Eostrobilops coreana
      • Eostrobilops kanjiokuboi
      • Eostrobilops humicolus
      • Eostrobilops diptyx
      • Eostrobilops taiwanica
      • Eostrobilops infrequens
      • Eostrobilops triptychus
    • Xuanchengospira flexilabrosa
    • Agallospira multispiralis
    • Palaeostrobilops antiquus
    • Coelostrobilops wenziana
  • Argnidae → (13 species)
    • Argna → (12 species)
      • Argna valsabina
      • Argna praeambula
      • Argna reperta
      • Argna bacillus
      • Argna bielzi
      • Argna biplicata
      • Argna jobae
      • Argna thracica
      • Argna ferrari
      • Argna bourguignatiana
      • Argna szekeresi
      • Argna pseudoennea
    • Speleodentorcula beroni
  • Orculidae → (83 species)
    • Orculella → (31 species)
      • Orculella astirakiensis
      • Orculella mayi
      • Orculella sirianocoriensis
      • Orculella nov
      • Orculella ignorata
      • Orculella connollyi
      • Orculella pfeiferi
      • Orculella cretilasithi
      • Orculella palatalis
      • Orculella garzanensis
      • Orculella menkhorsti
      • Orculella mesopotamica
      • Orculella exaggerata
      • Orculella creticostata
      • Orculella diensis
      • Orculella fodela
      • Orculella bulgarica
      • Orculella cretioreina
      • Orculella ruderalis
      • Orculella critica
      • Orculella garzaniensis
      • Orculella templorum
      • Orculella cretiminuta
      • Orculella aragonica
      • Orculella franciscoi
      • Orculella scalaris
      • Orculella creantirudis
      • Orculella cretimaxima
      • Orculella orientalis
      • Orculella heterostropha
      • Orculella pabsti
    • Sphyradium → (2 species)
      • Sphyradium doliolum
      • Sphyradium hasta
    • Schileykula → (13 species)
      • Schileykula inversa
      • Schileykula attilae
      • Schileykula maculata
      • Schileykula neuberti
      • Schileykula scyphus
      • Schileykula acampsis
      • Schileykula nordsiecki
      • Schileykula contraria
      • Schileykula robusta
      • Schileykula aculeata
      • Schileykula batumensis
      • Schileykula trapezensis
      • Schileykula sigma
    • Orcula → (32 species)
      • Orcula tomlini
      • Orcula pseudodolium
      • Orcula caabensis
      • Orcula mayi
      • Orcula austriaca
      • Orcula tolminensis
      • Orcula dolium
      • Orcula wolteri
      • Orcula farzughana
      • Orcula kaltenbachi
      • Orcula oviformis
      • Orcula klemmi
      • Orcula gregoryi
      • Orcula zilchi
      • Orcula alvarium
      • Orcula pabsti
      • Orcula connollyi
      • Orcula orientalis
      • Orcula conica
      • Orcula fuchsi
      • Orcula bisacchiae
      • Orcula farzughanajaeckeli
      • Orcula wagneri
      • Orcula gularis
      • Orcula nagaensis
      • Orcula spoliata
      • Orcula marmarica
      • Orcula jetschini
      • Orcula iraqensis
      • Orcula dobrogica
      • Orcula schmidtii
      • Orcula restituta
    • Pilorcula → (3 species)
      • Pilorcula trifilaris
      • Pilorcula aspinosa
      • Pilorcula pusilla
    • Alvariella multiplicata
    • Pupilorcula karaganica
  • Spelaeodiscidae → (22 species)
    • Spelaeodiscus → (11 species)
      • Spelaeodiscus latecostatus
      • Spelaeodiscus hauffeni
      • Spelaeodiscus triarius
      • Spelaeodiscus albanicus
      • Spelaeodiscus virpazarioides
      • Spelaeodiscus unidentatus
      • Spelaeodiscus dejongi
      • Spelaeodiscus obodensis
      • Spelaeodiscus densecostatus
      • Spelaeodiscus bulgaricus
      • Spelaeodiscus hunyadii
    • Virpazaria → (9 species)
      • Virpazaria aspectulabeatidis
      • Virpazaria adrianae
      • Virpazaria dhorai
      • Virpazaria nicoleae
      • Virpazaria ripkeni
      • Virpazaria pageti
      • Virpazaria stojaspali
      • Virpazaria backhuysi
      • Virpazaria deelemanorum
    • Klemmia → (2 species)
      • Klemmia magnicosta
      • Klemmia sinistrorsa
  • Pyramidulidae → (22 species)
    • Pyramidula solitaria
    • Pyramidula cephalonica
    • Pyramidula chorismenostoma
    • Pyramidula hierosolymitana
    • Pyramidula patagonica
    • Pyramidula gracilitesta
    • Pyramidula conica
    • Pyramidula recurrecta
    • Pyramidula umbilicata
    • Pyramidula alternata
    • Pyramidula shantungensis
    • Pyramidula erdli
    • Pyramidula capillata
    • Pyramidula schuppi
    • Pyramidula strigosa
    • Pyramidula saxatilis
    • Pyramidula kuznetsovi
    • Pyramidula mascallensis
    • Pyramidula javana
    • Rock snail (Pyramidula pusilla)
    • Pyramidula jaenensis
    • Rock snail (Pyramidula rupestris)
  • whorl snails (Vertiginidae) → (428 species)
    • Vertigininae → (226 species)
      • Vertigo → (146 species)
        • Pupa muscicola
        • Pupa diezi
        • Pupa minima
        • Pupa undorfensis
        • Isthmia dubia
        • Vertigo iowaensis
        • Vertigo arenula
        • Vertigo sieversi
        • cylindrical vertigo (Vertigo binneyana)
        • Vertigo sandbergeri
        • Vertigo gouldi
        • Pacific vertigo (Vertigo andrusiana)
        • Vertigo antipygmaea
        • Vertigo ultimathule
        • Vertigo whitei
        • capital vertigo (Vertigo oscariana)
        • threaded vertigo (Vertigo rowelli)
        • Vertigo illjeborgi
        • Vertigo arctica
        • Marsh Whorl Snail (Vertigo antivertigo)
        • deep-throat vertigo (Vertigo nylanderi)
        • cone cross vertigo (Vertigo occidentalis)
        • Vertigo nitens
        • Vertigo bifida
        • olive vertigo (Vertigo perryi)
        • Vertigo brusinai
        • Vertigo moenana
        • Vertigo arizonensis
        • bluff vertigo (Vertigo meramecensis)
        • Vertigo hachijoensis
        • ribbed california vertigo (Vertigo californica)
        • Wall Whorl Snail (Vertigo pusilla)
        • Vertigo heldi
        • crested vertigo (Vertigo pygmaea)
        • Vertigo circumlabiata
        • cross vertigo (Vertigo modesta)
        • Vertigo trolli
        • Vertigo elsheimensis
        • Vertigo japonica
        • Idaho vertigo (Vertigo idahoensis)
        • Vertigo pageti
        • five-tooth vertigo (Vertigo ventricosa)
        • Vertigo inserta
        • Vertigo consteniusi
        • small-tooth vertigo (Vertigo allyniana)
        • Vertigo maxillosa
        • Vertigo quinquedentata
        • Vertigo myrmido
        • Narrow-mouthed whorl snail (Vertigo angustior)
        • Vertigo turonica
        • Vertigo marciae
        • Vertigo torrei
        • chestnut vertigo (Vertigo sterkii)
        • Vertigo uncata
        • Vertigo praepusilla
        • Vertigo milleri
        • Vertigo praecoquis
        • Vertigo genesioides
        • Vertigo cupressicola
        • Vertigo congoensis
        • Vertigo defranceii
        • Vertigo ultima
        • Vertigo kodamai
        • Vertigo hohenstaufenorum
        • Des Moulin's Whorl Snail (Vertigo moulinsiana)
        • mitered vertigo (Vertigo concinnula)
        • Vertigo thaumasta
        • Vertigo chytryi
        • Vertigo microsphaera
        • ovate california vertigo (Vertigo berryi)
        • Vertigo roberti
        • Vertigo pisewensis
        • Vertigo pseudoantivertigo
        • heart vertigo (Vertigo hinkleyi)
        • Vertigo kurilensis
        • smallmouth vertigo (Vertigo parvula)
        • Vertigo pimuensis
        • Monte Sano vertigo (Vertigo wheeleri)
        • delicate vertigo (Vertigo bollesiana)
        • Vertigo atavuncula
        • Vertigo douvillei
        • palmetto vertigo (Vertigo oralis)
        • Vertigo diegoensis
        • Mountain Whorl Snail (Vertigo alpestris)
        • Vertigo globosa
        • Vertigo nitidula
        • blade vertigo (Vertigo milium)
        • Vertigo beringia
        • Vertigo ronnebyensis
        • Vertigo chiricahuensis
        • Vertigo globus
        • Keys vertigo (Vertigo hebardi)
        • Vertigo trinotata
        • six-whorl vertigo (Vertigo morsei)
        • Vertigo numellata
        • Vertigo micra
        • Vertigo kochi
        • Vertigo occulta
        • Vertigo botanicorum
        • Vertigo kroloppi
        • variable vertigo (Vertigo gouldii)
        • Vertigo nangaparbatensis
        • Vertigo corduensis
        • Vertigo hibbardi
        • Vertigo catalinaria
        • Lilljeborg's Whorl Snail (Vertigo lilljeborgi)
        • Vertigo extima
        • Vertigo moldavica
        • Vertigo superstriata
        • Vertigo geyeri
        • Vertigo parcedentata
        • Columbia vertigo (Vertigo columbiana)
        • Vertigo farquhari
        • Vertigo doliara
        • Vertigo oughtoni
        • Vertigo palangula
        • striate vertigo (Vertigo rugosula)
        • Vertigo brierensis
        • Vertigo pseudosubstriata
        • Vertigo kushiroensis
        • Vertigo pontileviensis
        • Vertigo bermudensis
        • Vertigo bandulana
        • horseshoe vertigo (Vertigo dalliana)
        • Vertigo hirasei
        • Vertigo tembrockae
        • Vertigo malleata
        • Callused Vertigo (Vertigo arthuri)
        • Striated Whorl Snail (Vertigo substriata)
        • cupped vertigo (Vertigo clappi)
        • Vertigo marki
        • Vertigo likharevi
        • Vertigo eogea
        • swamp vertigo (Vertigo teskeyae)
        • Vertigo griqualandica
        • Vertigo inflatula
        • honey vertigo (Vertigo tridentata)
        • Vertigo hydrophila
        • crested vertigo (Vertigo cristata)
        • crested vertigo (Vertigo genesii)
        • Vertigo saxicola
        • Alabama vertigo (Vertigo alabamensis)
        • Hanna's Vertigo (Vertigo hannai)
        • grassland whorl snail (Vertigo ovata)
        • tapered vertigo (Vertigo elatior)
        • Vertigo coloradensis
      • Afripupa → (10 species)
        • Afripupa bisulcata
        • Afripupa farquhari
        • Afripupa kanongae
        • Afripupa rodriguezensis
        • Afripupa pelengeae
        • Afripupa kenyensis
        • Afripupa griqualandica
        • Afripupa misaliensis
        • Afripupa vengoensis
        • Afripupa densestriata
      • Truncatellina → (56 species)
        • Truncatellina dysorata
        • Truncatellina cryptodonta
        • Truncatellina similis
        • Truncatellina haasi
        • Truncatellina cameroni
        • Truncatellina opisthodon
        • Truncatellina beckmanni
        • Truncatellina velkovrhi
        • Truncatellina upembae
        • Truncatellina uniarmata
        • Truncatellina insulivaga
        • Truncatellina quantula
        • Truncatellina belokrysi
        • Truncatellina silvicola
        • Truncatellina babusarica
        • Truncatellina sykesii
        • Truncatellina claustralis
        • Truncatellina ninagongonis
        • Truncatellina ayubiana
        • Truncatellina pygmaeorum
        • Truncatellina naivashaensis
        • Truncatellina algoviana
        • Truncatellina costulata
        • Cylindrical Whorl Snail (Truncatellina cylindrica)
        • Truncatellina biscoitoit
        • Truncatellina micra
        • Truncatellina obesa
        • Truncatellina callicratis
        • Truncatellina laeviuscula
        • Truncatellina portosantana
        • Truncatellina purpuraria
        • Truncatellina ruwenzoriensis
        • Truncatellina atomus
        • Truncatellina lussinensis
        • Truncatellina monodon
        • Truncatellina taurica
        • Truncatellina prainhana
        • Truncatellina biscoitoi
        • Truncatellina sundleri
        • Truncatellina maresquelli
        • Truncatellina bhutanensis
        • Truncatellina dentata
        • Truncatellina podolica
        • Truncatellina arboricola
        • Truncatellina flavogilva
        • Truncatellina pretoriensis
        • Truncatellina adami
        • Truncatellina blanfordi
        • Truncatellina strobeli
        • Truncatellina schilleri
        • Truncatellina rothi
        • Truncatellina iota
        • Truncatellina linearis
        • Truncatellina lardea
        • Truncatellina perplexa
        • Truncatellina subrothi
      • Columella → (14 species)
        • Columella hartmutnordsiecki
        • Columella pygmaeorum
        • Columella microspora
        • Columella nymphaepratensis
        • Columella aspera
        • Columella talgarica
        • mellow column (Columella columella)
        • Columella tridentata
        • Columella hasta
        • toothless column snail (Columella edentula)
        • Columella acicularis
        • Columella intermedia
        • high-spire column (Columella simplex)
        • Columella polvonense
    • Bothriopupa → (5 species)
      • Bothriopupa tenuidens
      • pitted birddrop (Bothriopupa variolosa)
      • Bothriopupa leucodon
      • Bothriopupa breviconus
      • Bothriopupa conoidea
    • Paraboysidia → (18 species)
      • Paraboysidia frequens
      • Paraboysidia kitteli
      • Paraboysidia paralella
      • Paraboysidia serpa
      • Paraboysidia hupeana
      • Paraboysidia wangviangensis
      • Paraboysidia laidlawi
      • Paraboysidia anguloobtusa
      • Paraboysidia boettgeri
      • Paraboysidia nabhitabhatai
      • Paraboysidia dohertyi
      • Paraboysidia tarutao
      • Paraboysidia freguens
      • Paraboysidia oreia
      • Paraboysidia tamphathai
      • Paraboysidia gittenbergeri
      • Paraboysidia pangmapaensis
      • Paraboysidia kelantanensis
    • Nesopupinae → (67 species)
      • Nesopupa → (55 species)
        • Nesopupa litoralis
        • Nesopupa disjuncta
        • Nesopupa moreleti
        • Nesopupa quadrasi
        • Nesopupa limatula
        • Nesopupa oahuensis
        • Nesopupa godeffroyi
        • Nesopupa tongana
        • Nesopupa minutalis
        • Nesopupa insularis
        • Nesopupa dispersa
        • Nesopupa anceyana
        • Nesopupa vinsoni
        • Nesopupa moluccana
        • Nesopupa dubitalis
        • Nesopupa mokaensis
        • Nesopupa multidentata
        • Nesopupa pleurophora
        • Nesopupa peilei
        • Nesopupa ponapica
        • Nesopupa scotti
        • Nesopupa maaseni
        • Nesopupa comorensis
        • Nesopupa novopommerana
        • Nesopupa angusta
        • Nesopupa lifouana
        • Nesopupa dentifera
        • Nesopupa hackerae
        • Nesopupa morini
        • Nesopupa subcentralis
        • Nesopupa gonioplax
        • Nesopupa waianaensis
        • Nesopupa ganzae
        • Nesopupa selebensis
        • Nesopupa tantilla
        • Nesopupa yamagutii
        • Nesopupa armata
        • Nesopupa malayana
        • Nesopupa tenimberica
        • Nesopupa madgei
        • A Terrestrial Snail (Nesopupa newcombi)
        • Nesopupa corrugata
        • Nesopupa paivae
        • Nesopupa rarotonga
        • Nesopupa cocosensis
        • Nesopupa mariei
        • Nesopupa moellendorffi
        • Nesopupa micra
        • Nesopupa tryphera
        • Nesopupa dubitabilis
        • Nesopupa galapagensis
        • Nesopupa singularis
        • Nesopupa nannodes
        • Nesopupa incerta
        • Nesopupa mooreana
      • Pronesopupa → (6 species)
        • Pronesopupa orycta
        • Pronesopupa boettgeri
        • Pronesopupa simplaria
        • Pronesopupa senex
        • Pronesopupa hystricella
        • Pronesopupa acanthinula
      • Negulus → (5 species)
        • Negulus reinhardti
        • Negulus amplinternatus
        • Negulus kenianus
        • Negulus abyssinicus
        • Negulus ruwenzoriensis
      • Staurodon seminulum
    • Gyliotrachela → (34 species)
      • Gyliotrachela muaklekensis
      • Gyliotrachela luctans
      • Gyliotrachela transitans
      • Gyliotrachela hungerfordiana
      • Gyliotrachela phoca
      • Gyliotrachela adela
      • Gyliotrachela khaochakan
      • Gyliotrachela srirachaensis
      • Gyliotrachela striolata
      • Gyliotrachela torticollis
      • Gyliotrachela frequens
      • Gyliotrachela plesiolopa
      • Gyliotrachela phupaman
      • Gyliotrachela kohrin
      • Gyliotrachela diarmaidi
      • Gyliotrachela depressispira
      • Gyliotrachela emergens
      • Gyliotrachela australis
      • Gyliotrachela crossei
      • Gyliotrachela troglodytes
      • Gyliotrachela saxicola
      • Gyliotrachela modesta
      • Gyliotrachela concreta
      • Gyliotrachela tridentatus
      • Gyliotrachela saraburiensis
      • Gyliotrachela fruhstorferi
      • Gyliotrachela khaochongensis
      • Gyliotrachela surakiti
      • Gyliotrachela muangon
      • Gyliotrachela burchi
      • Gyliotrachela dohertyi
      • Gyliotrachela khaowongensis
      • Gyliotrachela salpinx
      • Gyliotrachela khaowongkot
    • Sterkia → (7 species)
      • Sterkia calamitosa
      • Sterkia gittenbergeri
      • Sterkia bakeri
      • California birddrop (Sterkia hemphilli)
      • Sterkia antillensis
      • Caribbean birddrop (Sterkia eyriesii)
      • insular birddrop (Sterkia clementina)
    • Lyropupa → (21 species)
      • Lyropupa lyrata
      • Lyropupa anceyana
      • Lyropupa limatula
      • Lyropupa thaumasia
      • Lyropupa kahoolawensis
      • Lyropupa hawaiiensis
      • Lyropupa striatula
      • Lyropupa perlonga
      • Lyropupa micra
      • Lyropupa ovatula
      • Lyropupa kahoolavensis
      • Lyropupa spaldingi
      • Lyropupa ovotula
      • Lyropupa thaanumi
      • Lyropupa cyrta
      • Lyropupa costata
      • Lyropupa rhabdota
      • Lyropupa antiqua
      • Lyropupa magdalenae
      • Lyropupa mirabilis
      • Lyropupa prisca
    • Hypselostoma → (36 species)
      • Hypselostoma annamiticum
      • Hypselostoma pusillum
      • Hypselostoma rochebruni
      • Hypselostoma rupestre
      • Hypselostoma piconis
      • Hypselostoma satulensis
      • Hypselostoma everetti
      • Hypselostoma holimanae
      • Hypselostoma edentata
      • Hypselostoma roebeleni
      • Hypselostoma pattalungensis
      • Hypselostoma lacrima
      • Hypselostoma bensonianum
      • Hypselostoma khaowongensis
      • Hypselostoma chatnareeae
      • Hypselostoma erawan
      • Hypselostoma latispira
      • Hypselostoma quadrasi
      • Hypselostoma dayanum
      • Hypselostoma polyodon
      • Hypselostoma insularum
      • Hypselostoma panhai
      • Hypselostoma utongensis
      • Hypselostoma cucumensis
      • Hypselostoma edentulum
      • Hypselostoma loei
      • Hypselostoma benetuitum
      • Hypselostoma luzonicum
      • Hypselostoma kentingensis
      • Hypselostoma dilatatum
      • Hypselostoma cambodjense
      • Hypselostoma taehwani
      • Hypselostoma tubiferum
      • Hypselostoma discobasis
      • Hypselostoma socialis
      • Hypselostoma sibuyanicum
    • Nesoropupa → (4 species)
      • Nesoropupa fontainei
      • Nesoropupa duodecim
      • Nesoropupa nathaliae
      • Nesoropupa fenua
    • Glandicula → (2 species)
      • Glandicula gaali
      • Glandicula tiarula
    • Costigo → (3 species)
      • Costigo calamianica
      • Costigo saparuana
      • Costigo desmazuresi
    • Nearctula degeneris
    • Pseudelix pupula
    • Nesopuparia norfolkensis
    • Tetoripupa costata
    • Paracraticula umbra
  • Pleurodiscidae → (8 species)
    • Pleurodiscus → (7 species)
      • Pleurodiscus asteriscus
      • Pleurodiscus astericus
      • Pleurodiscus klemmi
      • Pleurodiscus sudensis
      • Pleurodiscus bellus
      • Pleurodiscus balmei
      • Pleurodiscus cyprius
    • Pleurodiscoides frici
  • chrysalis snails (Pupillidae) → (257 species)
    • Pupilla → (122 species)
      • Pupa intersecta
      • Pupa hoppii
      • Pupa tongriana
      • Pupa indigena
      • Pupa dalliana
      • Pupa heberti
      • Pupa antiquitata
      • Pupa arctica
      • Pupa coarctata
      • Pupa reussi
      • Pupa libanotica
      • Pupa trojana
      • Pupa diopsis
      • Pupa hebraica
      • Pupa incolata
      • Pupa elegans
      • Pupa phrygica
      • Pupa bisulcata
      • Pupa edentula
      • Pupa armifera
      • Pupa calcarea
      • Pupa turcica
      • Pupa palangula
      • Pupa vincta
      • Pupa subquadridens
      • Pupa mumiola
      • Pupa californica
      • Pupa hydrophila
      • Pupa amblymorpha
      • Pupa pretoriennis
      • Pupa bigsbii
      • Pupa fabrei
      • Pupa imbricata
      • Pupa psichion
      • Pupa spreta
      • Pupa helicoides
      • Pupa alticola
      • Pupa dohrniana
      • Pupa charybdia
      • Pupa turtoni
      • Pupa gorganica
      • Pupa canaliculata
      • Pupa bonneti
      • Pupa schweinfurthi
      • Pupa haploa
      • Pupa versipolis
      • Pupa inversa
      • Pupa palanga
      • Pupa striata
      • Pupa uvula
      • Pupa macilenta
      • Pupa cristata
      • Pupa fallax
      • Pupa capitani
      • Pupa socotrana
      • Pupilla blandii
      • Pupilla selecta
      • Moss chrysalis snail (Pupilla muscorum)
      • Pupilla afghanicum
      • Pupilla hudsoniana
      • Pupilla bigranata
      • Pupilla scapula
      • three-tooth column (Pupilla sonorana)
      • Pupilla bogdanovkaense
      • Pupilla sterrii
      • Pupilla chinensis
      • Pupilla aeoli
      • Pupilla satparanica
      • Pupilla triplicatoidea
      • Pupilla kyrostriata
      • Rocky Mountain column (Pupilla blandi)
      • Pupilla limata
      • Pupilla pratensis
      • Pupilla loessica
      • Pupilla paraturcmenica
      • Pupilla mlomnickii
      • Pupilla seminskii
      • Pupilla khunjerabica
      • Pupilla salemensis
      • Pupilla iheringi
      • Pupilla hokkaidoensis
      • Pupilla goniodon
      • Pupilla shantungensis
      • Pupilla zeisslerae
      • Pupilla sterkiana
      • Pupilla richthofeni
      • Pupilla gallae
      • Pupilla duplicata
      • Pupilla roberti
      • Pupilla inequidenta
      • Pupilla alluvionica
      • Pupilla perlabiata
      • Pupilla impressa
      • Pupilla hopeiensis
      • Pupilla tetrodus
      • Pupilla alabiella
      • Pupilla michaudi
      • Pupilla inops
      • Pupilla alpicola
      • Pupilla mutabilis
      • Pupilla inermis
      • crestless column (Pupilla hebes)
      • Pupilla triplicata
      • Pupilla altaica
      • Pupilla bargi
      • Pupilla signataeformis
      • Pupilla belokrysi
      • Pupilla australis
      • Pupilla desiderata
      • Pupilla parvula
      • Pupilla guadalupensis
      • Pupilla alaskensis
      • Pupilla bipatulata
      • Pupilla turcmenica
      • Pupilla ficulnea
      • Pupilla ziaratana
      • Pupilla grabaui
      • top-heavy column (Pupilla syngenes)
      • Pupilla anzobica
      • Pupilla striopolita
      • Pupilla cinghalensis
      • Pupilla staszicii
    • Anauchen → (16 species)
      • Anauchen srakeoensis
      • Anauchen huaykhakang
      • Anauchen chatnareeae
      • Anauchen khaochongpran
      • Anauchen utaithaniensis
      • Anauchen smokon
      • Anauchen banmiensis
      • Anauchen angulinus
      • Anauchen messageri
      • Anauchen informis
      • Anauchen chaunosalpinx
      • Anauchen sichang
      • Anauchen gereti
      • Anauchen chedi
      • Anauchen angthongensis
      • Anauchen depressus
    • Pupisoma → (31 species)
      • Pupisoma solemi
      • Pupisoma dioscoricola
      • Pupisoma mediamericanum
      • Pupisoma harpula
      • Pupisoma bilamellatum
      • Pupisoma japonicum
      • Pupisoma renschi
      • Pupisoma porti
      • Pupisoma dioscoricula
      • Pupisoma vermeuleni
      • Pupisoma longstaffi
      • Pupisoma orcula
      • Pupisoma costulata
      • Pupisoma hueense
      • Pupisoma evezardi
      • Gulf babybody (Pupisoma macneilli)
      • Pupisoma circumlitum
      • Pupisoma orcella
      • Pupisoma bailyi
      • Pupisoma lignicola
      • Pupisoma evada
      • Pupisoma chytrophora
      • Pupisoma waterloti
      • Pupisoma pagodula
      • Pupisoma umbilicata
      • Pupisoma comicolense
      • Pupisoma moleculina
      • Pupisoma misaliensis
      • Pupisoma mauritiana
      • Pupisoma unilamellatum
      • Pupisoma microturbinata
    • Microstele → (8 species)
      • Microstele noltei
      • Microstele mariae
      • Microstele muscerda
      • Microstele alamellata
      • Microstele huberi
      • Microstele wenzi
      • Microstele pingi
      • Microstele iredalei
    • Glyptopupoides → (2 species)
      • Glyptopupoides egregia
      • Pupoides xinyuensis
    • Gibbulinopsis → (8 species)
      • Gibbulinopsis gracilis
      • Gibbulinopsis pupula
      • Gibbulinopsis signata
      • Gibbulinopsis interrupta
      • Gibbulinopsis cryptodon
      • Gibbulinopsis nanosignata
      • Gibbulinopsis fontana
      • Gibbulinopsis rahtii
    • Pupoides → (35 species)
      • white-lip dagger (Pupoides albilabris)
      • Pupoides ischnus
      • Pupoides nitidulus
      • Pupoides adelaidae
      • Pupoides eupleura
      • Pupoides antiquus
      • Pupoides pacificus
      • Pupoides costatus
      • Pupoides chanlerensis
      • Pupoides orzolae
      • Pupoides lardeus
      • Pupoides gnocco
      • Pupoides tephrodes
      • Pupoides iredalei
      • Pupoides catalinensis
      • Pupoides montana
      • Pupoides guchengensis
      • Rocky Mountain dagger (Pupoides inornatus)
      • Pupoides lepidulus
      • Pupoides centralis
      • Pupoides taiwanicus
      • Pupoides beltianus
      • island dagger (Pupoides modicus)
      • Pupoides pilsbryi
      • Pupoides minimus
      • Pupoides cordatus
      • Pupoides myoporinae
      • Pupoides calaharicus
      • Pupoides seminulum
      • Pupoides coenopictus
      • ribbed dagger (Pupoides hordaceus)
      • Pupoides arizonensis
      • Pupoides contrarius
      • Pupoides eremicolus
      • Pupoides zhenjiangensis
    • Aulacospira → (15 species)
      • Aulacospira lampangensis
      • Aulacospira lens
      • Aulacospira triptychia
      • Aulacospira scalatella
      • Aulacospira khaopratun
      • Aulacospira conica
      • Aulacospira porrecta
      • Aulacospira hoholoma
      • Aulacospira pluangtong
      • Aulacospira mucronata
      • Aulacospira hololoma
      • Aulacospira krobyloides
      • Aulacospira smaesarnensis
      • Aulacospira rhombostoma
      • Aulacospira triptycha
    • Krobylos → (10 species)
      • Krobylos veruwan
      • Krobylos sinensis
      • Krobylos clerxi
      • Krobylos pomjuk
      • Krobylos maehongsonensis
      • Krobylos tampla
      • Krobylos maehungsonensis
      • Krobylos takensis
      • Krobylos laosensis
      • Krobylos kangkoy
    • Pumilicopta → (3 species)
      • Pumilicopta bifurcata
      • Pumilicopta solemorum
      • Pumilicopta kessneri
    • Cylindrovertilla → (4 species)
      • Cylindrovertilla kingi
      • Cylindrovertilla paitensis
      • Cylindrovertilla hedleyi
      • Cylindrovertilla fabreana
    • Prazoogenetes orcula
    • Albertanella minuta
    • Pupoidopsis hawaiensis
  • Spelaeoconcha paganettii

Other links

• Wikidata
• Wikipedia (language: en)
• Wikipedia (language: de)
• Wikipedia (language: it)
• Wikipedia (language: zh)
• Wikipedia (language: nl)
• Wikipedia (language: uk)
• Wikipedia (language: ko)
• Encyclopedia of Life
• Open Tree of Life

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