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Base of the evolutionary tree for dung beetles & tumblebugs (
Scarabaeinae
)
← parent
dung beetles & tumblebugs (
Scarabaeinae
) (1927 species)
Coprini
→
(286 species)
black dung beetles (
Copris
) →
(263 species)
Copris decellei
Copris youngai
Copris parapecuarius
Copris riekoae
Copris furciceps
Copris umbilicatus
Copris complexus
Copris fukiensis
Copris pueli
Copris jacchus
Copris jahi
Copris pecuarius
Copris davidi
Copris brevicornis
Copris andrewesi
Copris elphenor
Copris kachinensis
Copris vietnamicus
Copris denticulatus
Copris draco
Copris kasagii
Copris warneri
Copris fallax
Copris obenbergeri
Copris chimalapensis
Copris costaricensis
Copris pseudosinicus
Copris rebouchei
Copris klapperichi
Copris sphaeropterus
Copris cambeforti
Copris binodis
Mexican dung beetle (
Copris incertus
)
Copris truncatus
Copris neotridens
Copris subterraneus
Copris pseudomoffartsi
Copris antares
Copris uenoi
Copris surdus
Copris tridentatus
Copris susanae
Copris bengalensis
Copris vietnamimontanus
Copris armiger
Copris typhoeus
Copris cornifrons
Copris sexdentatus
Copris bacchus
Copris gibbulus
Copris megaceratoides
Copris szechouanicus
Copris costariensis
Copris tsukamotoi
Copris colmanti
Copris moffartsi
Copris keralensis
Copris mesacanthus
Copris ruricola
Copris poggii
Howden's Copris (
Copris howdeni
)
Copris zhangi
Copris lecontei
Copris interioris
Copris urus
Copris schoolmeestersi
Copris signatus
Copris anceus
Copris hatayamai
Copris lannathai
Copris orion
Copris obesus
Copris aeneus
Copris martinezi
Copris medogensis
Copris graueri
Copris caobangensis
Copris agnus
Copris gilleti
Copris jacchoides
Copris atropolitus
Copris fallaciosus
Copris boucardi
Copris cheni
Copris paraszechouanicus
Copris gladiator
Copris curlettii
Copris laius
Copris matthewsi
Copris freyi
Copris repertus
Copris phungae
Copris nevinsoni
Copris angustus
Copris potanini
Copris cribricollis
Copris hybridus
Copris sodalis
Copris subpunctatus
Copris lugubris
Copris martinae
Copris arcturus
Copris sinon
Copris angusticornis
Copris punctatus
Copris carmelita
Copris siamensis
Copris capensis
Copris laioides
Copris pseudobootes
Copris remotus
Copris maesi
Copris humilis
Copris subsidens
Copris vankhaii
Copris misellus
Copris eburneus
Copris minutus
Copris druidum
Copris bootes
Copris morphaeus
Copris coriarius
Copris illotus
Copris corpulentus
Copris tetraodon
Copris ribbei
Copris barclayi
Copris sorex
Copris vrydaghi
Copris elpheneroides
Copris nubilosus
Copris cambodiensis
Copris sacontala
Copris phylax
Copris armatus
Copris scorpio
Copris camerunus
Copris fidius
Copris kusakabei
Copris yangi
Copris pseudochus
Copris brittoni
Copris bovinus
Copris numa
Copris cassius
Copris sinicus
Copris arrowi
Copris pedarioides
Copris magicus
Copris ares
Copris diversus
Copris quasilaevigatus
Copris marcus
Copris laeviceps
Copris bellator
Copris caliginosus
Copris crassus
Copris punjabensis
Copris moroni
Copris hartli
Copris moechus
Copris aspericollis
Copris frankenbergerianus
Copris corniger
horned dung beetle (
Copris lunaris
)
Copris igualensis
Copris sarpedon
Copris kiuchii
Copris confucius
Copris armeniacus
Copris arimotoi
Copris ochus
Copris erratus
Copris sabinus
Copris kasanka
Copris afgoi
Copris pauliani
Copris macer
Copris delicatus
Copris davisoni
Copris tripartitus
Copris garambae
Copris anomiopseoides
Copris laiiformis
Copris ritsemae
Copris carinicus
Copris renwarti
Copris jucundus
Copris tortoise commensal scarab (
Copris gopheri
)
Copris macclevei
Copris siangensis
Copris hispanicus
Copris manni
Copris ciliatus
Copris amyntor
Copris dudleyi
Copris mongkhoni
Copris megasoma
Copris katangae
Copris simulator
Copris halffteri
Copris inhalatus
Copris imitans
Copris gazellarum
Copris cariniceps
Copris basipunctatus
Copris hispanus
Copris bidens
Copris felschei
Copris montivagus
Copris juanmorai
Copris vilhenai
Copris macacus
Copris orphanus
Copris leruei
Copris evanidus
Copris saundersi
Copris celebensis
Copris punctipennis
Copris spinator
Copris sumatrensis
Copris algol
Copris arizonensis
Copris ramosiceps
Copris nepos
Copris amabilis
Copris inemarginatus
A Scarab Beetle (
Copris fricator
)
Copris mexicanus
Copris caelatus
Copris novaki
Copris mourgliai
Copris integer
Copris davisi
Copris klugi
Copris laevigatus
Copris wiesei
Copris acutidens
Copris neglectus
Copris prevosti
Copris amazonicus
Copris victorini
Copris brachypterus
Copris cribratus
Copris simonettai
Copris sonensis
Copris puncticollis
Copris johkii
Copris insidiosus
Copris excisus
Copris inaequabilis
Copris iris
Copris gracilis
Copris basilewskyi
Copris subdolus
Copris serius
Copris dracunculus
Copris sierrensis
Copris singularis
Copris compressipennis
Copris usambaricus
Copris sallei
Copris rugosus
Coptodactyla
→
(16 species)
Coptodactyla glabricollis
Coptodactyla lesnei
Coptodactyla torresica
Coptodactyla matthewsi
Coptodactyla stereocera
Coptodactyla papua
Coptodactyla onitoides
Coptodactyla ducalis
Coptodactyla tuberculata
Coptodactyla brooksi
Coptodactyla nitida
Coptodactyla storeyi
Coptodactyla monstrosa
Coptodactyla depressa
Coptodactyla meridionalis
Coptodactyla merdeka
Litocopris
→
(4 species)
Litocopris simplex
Litocopris muticus
Litocopris punctiventris
Litocopris ruandanus
Homocopris
→
(3 species)
Homocopris buckleyi
Homocopris achamas
Homocopris torulosus
Sisyphini
→
(101 species)
Sisyphus
→
(69 species)
Sisyphus caffer
Sisyphus kibale
Sisyphus ethiopicus
Sisyphus goryi
Sisyphus araneolus
Sisyphus juheli
Sisyphus muricatus
Sisyphus alveatus
Sisyphus spinipes
Sisyphus minettii
Sisyphus ibaya
Sisyphus crispatus
Sisyphus granulifer
Sisyphus latus
Sisyphus submonticolus
Sisyphus bouyeri
Sisyphus genierorum
Sisyphus swazi
Sisyphus seminulum
Sisyphus cambeforti
Sisyphus jossoi
Sisyphus arboreus
Sisyphus laoticus
Sisyphus biarmatus
Sisyphus denticrus
Sisyphus mkomazi
Sisyphus umbraphilus
Sisyphus lobi
Sisyphus bicuariensis
Sisyphus thoracicus
Sisyphus africanus
Sisyphus fortuitus
Sisyphus nanniscus
Sisyphus walteri
Sisyphus sordidus
Sisyphus splendidus
Sisyphus felschei
Sisyphus struwei
Sisyphus inconspicuus
Sisyphus schaefferi
Sisyphus janssensi
Sisyphus auricomus
Sisyphus gazanus
Sisyphus werneri
Sisyphus neobornemisszanus
Sisyphus trichodichromicus
Sisyphus longipes
Sisyphus ocellatus
Sisyphus manni
Sisyphus costatus
Sisyphus moltaensis
Sisyphus bayanga
Sisyphus nodifer
Sisyphus faciculatus
Sisyphus neglectus
Sisyphus oralensis
Sisyphus desaegeri
Sisyphus perissinottoi
Sisyphus impressipennis
Sisyphus indicus
Sisyphus fasciculatus
Sisyphus mendicus
Sisyphus aages
Sisyphus morettoi
Sisyphus maniti
Sisyphus ongo
Sisyphus australis
Sisyphus popovi
Sisyphus usambaricus
Neosisyphus
→
(32 species)
Neosisyphus spinipes
Neosisyphus kuehni
Neosisyphus morenoi
Neosisyphus mirabilis
Neosisyphus tai
Neosisyphus drumonti
Neosisyphus jossoi
Neosisyphus negus
Neosisyphus astriatus
Neosisyphus moorei
Neosisyphus baoule
Neosisyphus macrorubrus
Neosisyphus tibialis
Neosisyphus werneri
Neosisyphus barbarossa
Neosisyphus setiger
Neosisyphus tembyi
Neosisyphus fortuitus
Neosisyphus bowringi
Neosisyphus calcaratus
Neosisyphus rubrus
Neosisyphus angulicollis
Neosisyphus basilewskyi
Neosisyphus morettoi
Neosisyphus ruber
Neosisyphus paschalidisae
Neosisyphus trochantericus
Neosisyphus quadricollis
Neosisyphus confrater
Neosisyphus armatus
Neosisyphus dierkensi
Neosisyphus tarantula
Ateuchini
→
(265 species)
Canthidium
→
(173 species)
Canthidium perceptibile
Canthidium kiesenwetteri
Canthidium viride
Canthidium guyanense
Canthidium nobile
Canthidium metallicum
Canthidium moroni
Canthidium haagi
Canthidium humerale
Canthidium lentum
Canthidium volxemi
Canthidium kelleri
Canthidium pseudopuncticolle
Canthidium flavicorne
Canthidium taurinum
Canthidium laetum
Canthidium elegantulum
Canthidium leucopterum
Canthidium gemmingeri
Canthidium magnum
Canthidium aeneolum
Canthidium foveolatum
Canthidium chabanaudi
Canthidium quercetorum
Canthidium bovinum
Canthidium hyla
Canthidium subdopuncticolle
Canthidium cognatum
Canthidium bituberifrons
Canthidium picipes
Canthidium annagabrielae
Canthidium tenebrosum
Canthidium viridicolle
Canthidium barbacenicum
Canthidium puncticolle
Canthidium riverai
Canthidium viridiobscurum
Canthidium atomarium
Canthidium luteum
Canthidium lebasi
Canthidium impressum
Canthidium macclevei
Canthidium basale
Canthidium flavipes
Canthidium globulum
Canthidium vespertinum
Canthidium titschacki
Canthidium flavum
Canthidium femoratum
Canthidium haroldi
Canthidium deyrollei
Canthidium breve
Canthidium pullus
Canthidium ardens
Canthidium erythropterum
Canthidium darwini
Canthidium puncticeps
Canthidium dispar
Canthidium sulcicolle
Canthidium funebre
Canthidium steinheili
Canthidium cereum
Canthidium sulcatum
Canthidium cupreum
Canthidium korschefskyi
Canthidium nitidum
Canthidium dimidiatum
Canthidium flabellatum
Canthidium decoratum
Canthidium rutilum
Canthidium histrio
Canthidium moestum
Canthidium glabricolle
Canthidium gracilipes
Canthidium violaceipenne
Canthidium moestrum
Canthidium marseuli
Canthidium multipunctatum
Canthidium seladon
Canthidium angulicolle
Canthidium latipleurum
Canthidium rufipes
Canthidium marianelae
Canthidium chrysis
Canthidium refulgens
Canthidium delgadoi
Canthidium quadridens
Canthidium marielae
Canthidium trinodosum
Canthidium andersoni
Canthidium monoceros
Canthidium caesareum
Canthidium melanocephalum
Canthidium prasinum
Canthidium atricolle
Canthidium opacum
Canthidium latum
Canthidium manni
Canthidium gigas
Canthidium variolosum
Canthidium pseudoperceptibile
Canthidium discopygidiale
Canthidium aurifex
Canthidium inerme
Canthidium nanum
Canthidium pseudaurifex
Canthidium punctatum
Canthidium persplendens
Canthidium semicupreum
Canthidium tuberifrons
Canthidium ovale
Canthidium dohrni
Canthidium sladeni
Canthidium ruficolle
Canthidium howdeni
Canthidium smithi
Canthidium imperiale
Canthidium basipunctatum
Canthidium stali
Canthidium depressum
Canthidium macroculare
Canthidium planovultum
Canthidium kraatzi
Canthidium coerulescens
Canthidium calidum
Canthidium epistomale
Canthidium orbiculatum
Canthidium angusticeps
Canthidium priscillae
Canthidium hespenheidei
Canthidium bokermanni
Canthidium clypeale
Canthidium convexifrons
Canthidium paranum
Canthidium guanacaste
Canthidium kirschi
Canthidium nigritum
Canthidium rufinum
Canthidium punctatostriatum
Canthidium atramentarium
Canthidium pallidoalatum
Canthidium cavifrons
Canthidium abbreviatum
Canthidium collare
Canthidium versicolor
Canthidium onthophagoides
Canthidium discolor
Canthidium escalerai
Canthidium onitoides
Canthidium aterrimum
Canthidium batesi
Canthidium miscellum
Canthidium parvulum
Canthidium gerstaeckeri
Canthidium bicolor
Canthidium cuprinum
Canthidium hypocrita
Canthidium aurichalceum
Canthidium auricolle
Canthidium splendidum
Canthidium inoptatum
Canthidium minimum
Canthidium thalassinum
Canthidium emoryi
Canthidium deplanatum
Canthidium centrale
Canthidium euchalceum
Canthidium laevigatum
Canthidium puncticollis
Canthidium tricolor
Canthidium margaritae
Canthidium obscurum
Canthidium lucidum
Trichillidium
→
(3 species)
Trichillidium quadridens
Trichillidium caingua
Trichillidium pilosum
Ateuchus
→
(88 species)
Ateuchus luciae
Ateuchus loricatus
Ateuchus confusus
Ateuchus freudei
Ateuchus halffteri
Ateuchus cujuchi
Ateuchus pauperatus
Ateuchus solisi
Ateuchus pygidialis
Ateuchus connexus
Ateuchus rispolii
Ateuchus hornai
Ateuchus ambiguus
Ateuchus pauki
Ateuchus fetteri
Ateuchus striatulus
Ateuchus laevicolle
Ateuchus scatimoides
Ateuchus floridensis
Ateuchus oblongus
Ateuchus hypocrita
Ateuchus fuscorubrus
Ateuchus fuscipes
Ateuchus semicribratus
Ateuchus tridenticeps
Ateuchus subquadratus
Ateuchus laetitiae
Ateuchus hoplopygus
Ateuchus viduus
Ateuchus zoebischi
Ateuchus granigerus
Ateuchus gershensoni
Ateuchus contractus
Ateuchus punctatissimum
Ateuchus aeneomicans
Ateuchus howdeni
Ateuchus columbianus
Ateuchus alvarezi
Ateuchus pius
Ateuchus opacipennis
Ateuchus carcavalloi
Ateuchus femoratus
Ateuchus chrysopyge
Ateuchus alutacius
Ateuchus perezvelai
Ateuchus tuza
Ateuchus calcaratus
Ateuchus myrmecophilus
Ateuchus hamatus
Ateuchus breve
Ateuchus hendrichsi
Ateuchus balthasari
Ateuchus parvus
Ateuchus carolinae
Ateuchus globulus
Ateuchus perpusillus
Ateuchus robustus
Ateuchus nitidulus
Ateuchus procerus
Ateuchus floridanus
Ateuchus spinipes
Ateuchus viduum
Ateuchus steinbachi
Ateuchus vigilans
Ateuchus pruneus
Ateuchus mutilatus
Ateuchus ecuadorensis
Ateuchus vividus
Ateuchus simplex
Ateuchus colossus
Ateuchus apicatus
Ateuchus romani
Ateuchus bordoni
Ateuchus asperatus
Ateuchus illaesus
Ateuchus earthorus
Ateuchus texanus
Ateuchus alipioi
Ateuchus semicupreum
Ateuchus rodriguezi
Ateuchus histeroides
Ateuchus ginae
Ateuchus frontale
Ateuchus laterale
Ateuchus squalidus
Ateuchus benitojuarezi
Ateuchus ecuadorense
Ateuchus viridimicans
Leotrichillum louzadaorum
Gymnopleurini
→
(116 species)
Gymnopleurus
→
(60 species)
Gymnopleurus puncticollis
Gymnopleurus moerens
Gymnopleurus tristis
Gymnopleurus particolor
Gymnopleurus colmanti
Gymnopleurus ruandensis
Gymnopleurus aeruginosus
Gymnopleurus naviauxi
Gymnopleurus laevicollis
Gymnopleurus reichei
Gymnopleurus hypocrita
Gymnopleurus pustulatus
Gymnopleurus atratus
Gymnopleurus coerulescens
Gymnopleurus ignitus
Gymnopleurus aciculatus
Gymnopleurus parvus
Gymnopleurus koenigi
Gymnopleurus tuxeni
Gymnopleurus geoffroyi
Gymnopleurus elegans
Gymnopleurus deperditus
Gymnopleurus nyankpalaensis
Gymnopleurus gemmatus
Gymnopleurus aenescens
Gymnopleurus mimus
Gymnopleurus plicatus
Gymnopleurus mopsus
Gymnopleurus plicatulus
Gymnopleurus leei
Gymnopleurus foricarius
Gymnopleurus bicallosus
Gymnopleurus asperrimus
Gymnopleurus bicolor
Gymnopleurus sericeifrons
Gymnopleurus lacunosus
Gymnopleurus sindensis
Gymnopleurus bombayensis
Gymnopleurus fulgidus
Gymnopleurus flagellatus
Gymnopleurus latreillei
Gymnopleurus imitator
Gymnopleurus andreaei
Gymnopleurus somaliensis
Gymnopleurus biharensis
Gymnopleurus virens
Gymnopleurus purpureus
Gymnopleurus rhodesianus
Gymnopleurus qurosh
Gymnopleurus humeralis
Gymnopleurus miliaris
Gymnopleurus persianus
Gymnopleurus cyaneus
Gymnopleurus eocaenicus
Gymnopleurus jacksoni
Gymnopleurus profanus
Gymnopleurus thelwalli
Gymnopleurus sturmi
Gymnopleurus thoracicus
Gymnopleurus humanus
Allogymnopleurus
→
(19 species)
Allogymnopleurus alluaudi
Allogymnopleurus sericeicollis
Allogymnopleurus youngai
Allogymnopleurus anthracinus
Allogymnopleurus olivieri
Allogymnopleurus umbrinus
Allogymnopleurus chloris
Allogymnopleurus splendidus
Allogymnopleurus histrio
Allogymnopleurus consocius
Allogymnopleurus infranitens
Allogymnopleurus signaticollis
Allogymnopleurus maculosus
Allogymnopleurus thalassinus
Allogymnopleurus jeanneli
Allogymnopleurus aeneus
Allogymnopleurus spilotus
Allogymnopleurus sagnai
Allogymnopleurus indigaceus
Garreta
→
(26 species)
Garreta lugens
Garreta azareus
Garreta australugens
Garreta sumptuosus
Garreta namalugens
Garreta fastiditus
Garreta laetus
Garreta bechynei
Garreta malleolus
Garreta sylvestris
Garreta ruficornis
Garreta smaragdifer
Garreta caffer
Garreta unicolor
Garreta gilleti
Garreta diffinis
Garreta nitens
Garreta mundus
Garreta dejeani
Garreta mombelgi
Garreta wahlbergi
Garreta opacus
Garreta matabelensis
Garreta morosus
Garreta nyassicus
Garreta crenulatus
Paragymnopleurus
→
(11 species)
Paragymnopleurus brahminus
Paragymnopleurus striatus
Paragymnopleurus sinuatus
Paragymnopleurus martinezi
Paragymnopleurus spinotus
Paragymnopleurus melanarius
Paragymnopleurus sparsus
Paragymnopleurus maurus
Paragymnopleurus ambiguus
Paragymnopleurus sumatrensis
Paragymnopleurus stipes
Australasian clade
→
(95 species)
Lepanus
→
(27 species)
Lepanus storeyi
Lepanus gressitti
Lepanus pygmaeus
Lepanus latheticus
Lepanus gelasinus
Lepanus palumensis
Lepanus nitidus
Lepanus occidentalis
Lepanus bournei
Lepanus dichrous
Lepanus ustulatus
Lepanus illawarensis
Lepanus bidentatus
Lepanus villosus
Lepanus howdeni
Lepanus politus
Lepanus pisoniae
Lepanus ovatus
Lepanus furcifer
Lepanus globulus
Lepanus fossulatus
Lepanus nr. pygmaeus COL2392
Lepanus vestitus
Lepanus monteithi
Lepanus australis
Lepanus parapisoniae
Lepanus papuanus
Matthewsius
→
(6 species)
Matthewsius penelopae
Matthewsius vanleeuweni
Matthewsius speewah
Matthewsius illawarrensis
Matthewsius stricklandensis
Matthewsius rossi
Amphistomus
→
(24 species)
Amphistomus pygmaeus
Amphistomus accidatus
Amphistomus cunninghamensis
Amphistomus speculatus
Amphistomus storeyi
Amphistomus macphersonensis
Amphistomus speculifer
Amphistomus inermis
Amphistomus calcaratus
Amphistomus trispiculatus
Amphistomus alfurorum
Amphistomus monticola
Amphistomus opacus
Amphistomus pectoralis
Amphistomus kukali
Amphistomus primonactus
Amphistomus fasciculatus
Amphistomus respiciens
Amphistomus complanatus
Amphistomus palpebratus
Amphistomus uncinatus
Amphistomus montanus
Amphistomus squalidus
Amphistomus tuberosus
Saphobius
→
(15 species)
Saphobius wakefieldi
Saphobius inflatipes
Saphobius curvipes
Saphobius squamulosus
Saphobius tibialis
Saphobius lesnei
Saphobius fulvipes
Saphobius laticollis
Saphobius edwardsi
Saphobius lepidus
Saphobius arrowi
Saphobius setosus
Saphobius fuscus
Saphobius brouni
Saphobius nitidulus
Canthonosoma
→
(3 species)
Canthonosoma mastersi
Canthonosoma castelnaui
Canthonosoma macleayi
Sauvagesinella
→
(4 species)
Sauvagesinella palustris
Sauvagesinella monstrosa
Sauvagesinella loftyensis
Sauvagesinella becki
Cephalodesmius
→
(3 species)
Cephalodesmius laticollis
Cephalodesmius quadridens
Cephalodesmius armiger
Aptenocanthon
→
(6 species)
Aptenocanthon winyar
Aptenocanthon wollumbin
Aptenocanthon jimara
Aptenocanthon kabura
Aptenocanthon hopsoni
Aptenocanthon monteithi
Monteithocanthon
→
(6 species)
Monteithocanthon arator
Monteithocanthon elliotensis
Monteithocanthon koombooloomba
Monteithocanthon paraarator
Monteithocanthon peckorum
Monteithocanthon glaber
Coproecus hemisphaericus
Eucraniini
→
(15 species)
Eucranium
→
(6 species)
Eucranium belenae
Eucranium simplicifrons
Eucranium arachnoides
Eucranium planicolle
Eucranium cyclosoma
Eucranium dentifrons
Anomiopsoides
→
(5 species)
Anomiopsoides heteroclyta
Anomiopsoides xerophila
Anomiopsoides fedemariai
Anomiopsoides biloba
Anomiopsoides cavifrons
Glyphoderus
→
(3 species)
Glyphoderus monticola
Glyphoderus centralis
Glyphoderus sterquilinus
Ennearabdus lobocephalus
Deltochilini
→
(818 species)
Epirinus
→
(30 species)
Epirinus montanus
Epirinus pygidialis
Epirinus sebastiani
Epirinus rugosus
Epirinus aeneus
Epirinus pseudorugosus
Epirinus asper
Epirinus convexus
Epirinus flagellatus
Epirinus relictus
Epirinus scrobiculatus
Epirinus validus
Epirinus obtusus
Epirinus hopei
Epirinus minimus
Epirinus gratus
Epirinus bentoi
Epirinus punctatus
Epirinus mucrodentatus
Epirinus striatus
Epirinus aquilus
Epirinus ngomae
Epirinus hilaris
Epirinus granulatus
Epirinus hluhluwensis
Epirinus davisi
Epirinus sulcipennis
Epirinus comosus
Epirinus drakomontanus
Epirinus silvestris
Diorygopyx
→
(8 species)
Diorygopyx simpliciclunis
Diorygopyx incrassatus
Diorygopyx incomptus
Diorygopyx tibialis
Diorygopyx niger
Diorygopyx cuspidatus
Diorygopyx asciculifer
Diorygopyx duplodentatus
Odontoloma
→
(20 species)
Odontoloma doubei
Odontoloma louwi
Odontoloma obscurum
Odontoloma spinicaudum
Odontoloma quadridens
Odontoloma cambeforti
Odontoloma disalatum
Odontoloma multipunctatum
Odontoloma endroedyi
Odontoloma planatum
Odontoloma pauliani
Odontoloma pygidiale
Odontoloma peckorum
Odontoloma metasternale
Odontoloma dentinum
Odontoloma sculpturatum
Odontoloma relictum
Odontoloma apiculum
Odontoloma pauxillum
Odontoloma pusillum
Epactoides
→
(37 species)
Epactoides masoalae
Epactoides humberti
Epactoides spinicollis
Epactoides mangabeensis
Epactoides costatus
Epactoides ankasokae
Epactoides perinetanus
Epactoides mahaboi
Epactoides olsoufieffi
Epactoides frontalis
Epactoides sambavae
Epactoides vadoni
Epactoides semiaeneus
Epactoides fiorii
Epactoides mananarae
Epactoides betschi
Epactoides hyphydroides
Epactoides jounii
Epactoides emkeni
Epactoides lissus
Epactoides vondrozoi
Epactoides major
Epactoides femoralis
Epactoides helenae
Epactoides andringitrae
Epactoides madecassus
Epactoides viridicollis
Epactoides vagecarinatus
Epactoides lacustris
Epactoides rahagai
Epactoides pauliani
Epactoides perrieri
Epactoides incertus
Epactoides tiinae
Epactoides basilewskyi
Epactoides hanskii
Epactoides paradoxus
Gyronotus
→
(11 species)
Marecon rufipennis
Gyronotus pumilus
Gyronotus dispar
Gyronotus fimetarius
Gyronotus marginatus
Gyronotus mulanjensis
Gyronotus perissinottoi
Gyronotus glabrosus
Gyronotus carinatus
Gyronotus rufipennis
Gyronotus schuelei
Peckolus
→
(3 species)
Peckolus parvus
Peckolus poenskopius
Peckolus alpinus
Canthonella
→
(16 species)
Canthonella constans
Canthonella gomezi
Canthonella instriata
Canthonella catharinensis
Canthonella baorucensis
Canthonella jarmilae
Canthonella howdeni
Canthonella quesqueya
Canthonella leptoculata
Canthonella barrerai
Canthonella parva
Canthonella silphoides
Canthonella isabellae
Canthonella sikesi
Canthonella amazonica
Canthonella lenkoi
Monoplistes
→
(6 species)
Monoplistes haroldi
Monoplistes curvipes
Monoplistes tropicus
Monoplistes leai
Monoplistes phanophilus
Monoplistes occidentalis
Aphengoecus
→
(2 species)
Aphengoecus multiserratus
Aphengoecus clypeatus
Namakwanus
→
(5 species)
Namakwanus minutus
Namakwanus davisi
Namakwanus scholtzi
Namakwanus irishi
Namakwanus kamfferi
Paracanthon
→
(4 species)
Paracanthon trichonotulus
Paracanthon hirsutus
Paracanthon rosinae
Paracanthon pereirai
Temnoplectron
→
(21 species)
Temnoplectron diversicolle
Temnoplectron involucre
Temnoplectron cooki
Temnoplectron finnigani
Temnoplectron laeve
Temnoplectron major
Temnoplectron aeneopiceum
Temnoplectron boucomonti
Temnoplectron waigeoensis
Temnoplectron monteithi
Temnoplectron reyi
Temnoplectron rotundum
Temnoplectron aeneolum
Temnoplectron bornemisszai
Temnoplectron subvolitans
Temnoplectron lewisense
Temnoplectron politulum
Temnoplectron wareo
Temnoplectron disruptum
Temnoplectron arfakensis
Temnoplectron atropolitum
Byrrhidium
→
(2 species)
Byrrhidium convexum
Byrrhidium namaquensis
Deltochilum
→
(107 species)
Deltochilum valgum
Deltochilum bordoni
Humpback Dung Beetle (
Deltochilum gibbosum
)
Deltochilum streblopodum
Deltochilum tessellatum
Deltochilum furcatum
Deltochilum arrowi
Deltochilum batesi
Deltochilum multicolor
Deltochilum spinipes
Deltochilum silphoides
Deltochilum viridicatum
Deltochilum cupreicolle
Deltochilum abdominalis
Deltochilum acanthus
Deltochilum rosamariae
Deltochilum carrilloi
Deltochilum tumidum
Deltochilum barbipes
Deltochilum speciosissimum
Deltochilum amazonicum
Deltochilum mexicanum
Deltochilum pseudoicarus
Deltochilum kolleri
Deltochilum feeri
Deltochilum finestriatum
Deltochilum punctatum
Deltochilum luederwaldti
Deltochilum elevatum
Deltochilum sextuberculatum
Deltochilum gigante
Deltochilum femorale
Deltochilum titovidaurrei
Deltochilum verruciferum
Deltochilum riehli
Deltochilum irroratum
Deltochilum diringshofeni
Deltochilum mourei
Deltochilum icaroides
Deltochilum lobipes
Deltochilum eurymedon
Deltochilum louzadai
Deltochilum paresi
Deltochilum sculpturatum
Deltochilum erodioides
Deltochilum violetae
Deltochilum larseni
Deltochilum pretiosum
Deltochilum aureopilosum
Deltochilum brasiliense
Deltochilum robustus
Deltochilum icariforme
Deltochilum subrubrum
Deltochilum septemstriatum
Deltochilum parile
Deltochilum morbillosum
Deltochilum scabriusculum
Deltochilum laetiusculum
Deltochilum aspericolle
Deltochilum granulosum
Deltochilum icarus
Deltochilum aequinoctiale
Deltochilum pseudoparile
Deltochilum peruanum
Deltochilum submetallicum
Deltochilum longiceps
Deltochilum fuscocupreum
Deltochilum arturoi
Deltochilum orbignyi
Deltochilum calcaratum
Deltochilum acropyge
Deltochilum molanoi
Deltochilum ritamourae
Deltochilum lindemannae
Deltochilum panamensis
Deltochilum carinatum
Deltochilum inaequale
Deltochilum violaceum
Deltochilum crenulipes
Deltochilum orbiculare
Deltochilum bezdeki
Deltochilum cristinae
Deltochilum viridescens
Deltochilum variolosum
Deltochilum enceladus
Deltochilum elongatum
Deltochilum alpercata
Deltochilum komareki
Deltochilum guildingii
Deltochilum rubripenne
Deltochilum cristatum
Deltochilum granulatum
Deltochilum mariafernandae
Deltochilum costalimai
Deltochilum guyanense
Deltochilum howdeni
Deltochilum viridicupreum
Deltochilum hypponum
Deltochilum kolbei
Deltochilum trisignatum
Deltochilum schefflerorum
Deltochilum dentipes
Deltochilum amandaarcanjoae
Deltochilum cangalha
Deltochilum sericeum
Deltochilum plebejum
Deltochilum loperae
Pseudonthobium
→
(9 species)
Pseudonthobium foveolatum
Pseudonthobium fracticolloides
Pseudonthobium humboldtense
Pseudonthobium paniense
Pseudonthobium doense
Pseudonthobium fracticolle
Pseudonthobium monteithianum
Pseudonthobium sinuatotibiale
Pseudonthobium genieri
Scybalocanthon
→
(20 species)
Scybalocanthon balachowskyi
Scybalocanthon trimaculatus
Scybalocanthon pygidialis
Scybalocanthon sexspilotus
Scybalocanthon nigellus
Scybalocanthon imitans
Scybalocanthon arcabuquensis
Scybalocanthon kelleri
Scybalocanthon kaestneri
Scybalocanthon aereus
Scybalocanthon moniliatus
Scybalocanthon cyanocephalus
Scybalocanthon nigriceps
Scybalocanthon darlingtoni
Scybalocanthon korasakiae
Scybalocanthon pinopterus
Scybalocanthon batesi
Scybalocanthon magnus
Scybalocanthon maculatus
Scybalocanthon zischkai
Malagoniella
→
(10 species)
Malagoniella astyanax
Malagoniella coerulea
Malagoniella chalybaea
Malagoniella bicolor
Malagoniella lanei
Malagoniella cupreicollis
Malagoniella puncticollis
Malagoniella magnifica
Malagoniella argentina
Malagoniella virens
Hansreia
→
(6 species)
Hansreia coriacea
Hansreia affinis
Hansreia oxygona
Hansreia grossii
Hansreia peugeoti
Hansreia krinskii
Scybalophagus
→
(4 species)
Scybalophagus plicatipennis
Scybalophagus patagonicus
Scybalophagus rugosus
Scybalophagus pilosus
Bohepilissus
→
(2 species)
Bohepilissus subtilis
Bohepilissus nitidus
Cryptocanthon
→
(40 species)
Cryptocanthon denticulum
Cryptocanthon otonga
Cryptocanthon cristobalensis
Cryptocanthon lindemanae
Cryptocanthon medinae
Cryptocanthon montebello
Cryptocanthon paradoxus
Cryptocanthon gilli
Cryptocanthon altus
Cryptocanthon humidus
Cryptocanthon genieri
Cryptocanthon buriticaorum
Cryptocanthon chiriquinus
Cryptocanthon howdeni
Cryptocanthon newtoni
Cryptocanthon andersoni
Cryptocanthon denticulatum
Cryptocanthon campbellorum
Cryptocanthon osaensis
Cryptocanthon foveatus
Cryptocanthon punctatus
Cryptocanthon curticrinis
Cryptocanthon pulidoae
Cryptocanthon escobari
Cryptocanthon urguensis
Cryptocanthon pumilus
Cryptocanthon lobipygus
Cryptocanthon ocosingo
Cryptocanthon bochilae
Cryptocanthon solisi
Cryptocanthon brevisetosus
Cryptocanthon pulidae
Cryptocanthon galbao
Cryptocanthon nebulinus
Cryptocanthon lobatus
Cryptocanthon napoensis
Cryptocanthon parvus
Cryptocanthon peckorum
Cryptocanthon andradei
Cryptocanthon rayonensis
Panelus
→
(34 species)
Panelus jaccoudi
Panelus watanabei
Panelus manasi
Panelus fallax
Panelus rufulus
Panelus setosus
Panelus orousseti
Panelus kalimantanicus
Panelus brendelli
Panelus rhodesiensis
Panelus besucheti
Panelus danumensis
Panelus borneensis
Panelus kubotai
Panelus mussardi
Panelus keralai
Panelus pernitidus
Panelus sabahi
Panelus mendeli
Panelus assamensis
Panelus deuvei
Panelus manmiaoae
Panelus ceylonicus
Panelus saigusai
Panelus tonkinensis
Panelus maedai
Panelus ovatus
Panelus crenatus
Panelus imitator
Panelus bakeri
Panelus parvulus
Panelus fujitai
Panelus wangi
Panelus puncticollis
Nanos
→
(41 species)
Nanos sinuatipes
Nanos pseudofusconitens
Nanos antsalovaensis
Nanos occidentalis
Nanos minutus
Nanos andreiae
Nanos humeralis
Nanos pseudominutus
Nanos mixtus
Nanos mirjae
Nanos incertus
Nanos bemarahaensis
Nanos ankaranae
Nanos viettei
Nanos semiscribosus
Nanos fusconitens
Nanos dubitatus
Nanos magnus
Nanos pygmaeus
Nanos nitens
Nanos antsihanakensis
Nanos pseudorubromaculatus
Nanos ater
Nanos bimaculatus
Nanos binotatus
Nanos punctatus
Nanos ranomafanaensis
Nanos manomboensis
Nanos peyrierasi
Nanos humbloti
Nanos hanskii
Nanos viridissimus
Nanos constricticollis
Nanos marojejyensis
Nanos vadoni
Nanos bicoloratus
Nanos neoelectrinus
Nanos clypeatus
Nanos rubrosignatus
Nanos pseudoviettei
Nanos manongorivoensis
Anachalcos
→
(2 species)
Anachalcos convexus
Anachalcos suturalis
Onthobium
→
(14 species)
Onthobium kuscheli
Onthobium montrouzieri
Onthobium lerati
Onthobium howdeni
Onthobium cooki
Onthobium gutierrezi
Onthobium macleayi
Onthobium boucheti
Onthobium tibialoides
Onthobium montheiti
Onthobium paniei
Onthobium tillieri
Onthobium asperatum
Onthobium cookii
Tumblebugs (
Canthon
) →
(171 species)
Canthon montanus
Canthon coerulescens
Canthon candens
Canthon curvodilatatus
Canthon substriatus
Canthon lecontei
A Scarab Beetle (
Canthon viridis
)
Canthon imitator
Canthon politus
Canthon corpulentus
Canthon aequinoctialis
Canthon septemmaculatus
Canthon delgadoi
Canthon sulcatus
Canthon edentulus
Canthon pallidus
Canthon smaragdulus
Canthon moniliatus
Canthon fallax
Canthon chiriguano
Canthon humboldti
Canthon marmoratus
Canthon octodentatus
Canthon subcyaneus
Canthon bisignatus
Canthon occidentalis
Canthon variabilis
Canthon hartmanni
Canthon callosus
Canthon ornatus
Canthon incisus
Canthon obscuriellus
Canthon tetraodon
Canthon xanthopus
Canthon corporaali
Canthon unicolor
Canthon circulatus
Canthon lunatus
Canthon antoniomartinezi
Canthon subhyalinus
Canthon caelius
Canthon viduus
Canthon ohausi
Canthon dentiger
Canthon planus
Canthon edmondsi
Canthon coloratus
Canthon formosus
Canthon pacificus
Canthon heyrovskyi
Canthon angularis
Canthon latipes
Canthon blumei
Canthon humectus
Canthon moroni
Canthon brunnipennis
Canthon oliverioi
Canthon juvencus
Canthon orfilai
Canthon plagiatus
Canthon fuscipes
Canthon bicolor
Canthon obliquus
Canthon depressipennis
Canthon quadratus
Canthon deyrollei
Canthon meridionalis
Canthon rzedowskii
Canthon rutilans
Canthon curvipes
Canthon deplanatus
Canthon nigripennis
Canthon dives
Canthon histrio
Canthon puncticollis
Canthon deloyai
Canthon helleri
Canthon manantlanensis
Canthon podagricus
Canthon championi
Canthon cyanellus
Canthon muticus
Canthon bispinus
Canthon mutabilis
Canthon quinquemaculatus
Canthon virens
Canthon carbonarius
Canthon piluliformis
Canthon violaceus
Canthon leechi
Canthon delicatulus
Canthon integricollis
Canthon melancholicus
Canthon indigaceus
Canthon honsi
Canthon ebenus
Canthon semiopacus
Canthon forcipatus
Canthon aberrans
Canthon perseverans
Canthon triangularis
Canthon ateuchiceps
Canthon velutinus
Canthon coahuilensis
Canthon inusitatus
Canthon probus
Canthon fulgidus
Canthon principalis
Canthon lucreciae
Canthon femoralis
Canthon laminatus
Canthon reyesi
Canthon simplex
Canthon daguerrei
Canthon zuninoi
Canthon forreri
Canthon steinheili
Canthon corruscus
Canthon acutiformis
Canthon columbianus
A Tumbling Dung Beetle (
Canthon praticola
)
A Scarab Beetle (
Canthon vigilans
)
Canthon perplexus
Canthon pilularius
Canthon denticulatus
Canthon lividus
Canthon delpontei
A Scarab Beetle (
Canthon chalcites
)
Canthon lamprimus
Canthon modestus
Canthon ebeneus
Canthon cobosi
Canthon lituratus
Canthon ibarragrassoi
Canthon acutus
Canthon janthinus
Canthon rubrescens
Canthon nyctelius
Canthon morsei
Canthon splendidus
Canthon reichei
Canthon simulans
Canthon cinctellus
Canthon coeruleicollis
Canthon bimaculatus
Canthon quadriguttatus
Canthon melanus
Canthon vulcanoae
Canthon signifer
Canthon mixtus
Canthon luteicollis
Canthon maldonadoi
Canthon brunneus
Canthon quadripunctatus
Canthon silvaticus
Canthon vazquezae
Canthon lafargei
Canthon angustatus
Canthon auricollis
Canthon riverai
Canthon matthewsi
Canthon monilifer
Canthon doesburgi
Canthon balteatus
Canthon gemellatus
Canthon sordidus
Canthon acutoides
Canthon staigi
Canthon sericatus
Canthon divinator
Canthon euryscelis
Tesserodoniella
→
(2 species)
Tesserodoniella meridionalis
Tesserodoniella elguetai
Pseudignambia
→
(2 species)
Pseudignambia mimerops
Pseudignambia squamata
Boletoscapter
→
(2 species)
Boletoscapter cornutus
Boletoscapter furcatus
Haroldius
→
(37 species)
Haroldius stevensi
Haroldius oharai
Haroldius rugatulus
Haroldius cardoni
Haroldius modestus
Haroldius brendelli
Haroldius discoidalis
Haroldius hwangi
Haroldius convexus
Haroldius leleupi
Haroldius lassallei
Haroldius fairmairei
Haroldius maruyamai
Haroldius turnai
Haroldius fleutiauxi
Haroldius sumatranus
Haroldius jechaer
Haroldius ennearthrus
Haroldius thailandensis
Haroldius stefanii
Haroldius kawadai
Haroldius kolaka
Haroldius pahangensis
Haroldius tangkoko
Haroldius loebli
Haroldius heimi
Haroldius philippinensis
Haroldius cambeforti
Haroldius uenoi
Haroldius annandalei
Haroldius borneensis
Haroldius globosus
Haroldius perroti
Haroldius celebensis
Haroldius pauliani
Haroldius herrenorum
Haroldius penelopae
Ignambia
→
(2 species)
Ignambia fasciculata
Ignambia fascicularis
Megathopa
→
(2 species)
Megathopa villosa
Megathopa violacea
Arachnodes
→
(93 species)
Epilissus genieri
Epilissus guillaumeti
Epilissus cuprarius
Epilissus striatoides
Epilissus antoetrae
Epilissus sikorai
Epilissus emmae
Epilissus fuscovirens
Epilissus morio
Epilissus armipes
Epilissus apotolamproides
Epilissus splendidus
Epilissus prasinus
Epilissus alluaudi
Epilissus fantamattii
Epilissus niger
Epilissus planatus
Epilissus ruteri
Epilissus viridis
Epilissus andranobeensis
Arachnodes sikorai
Arachnodes cyanescens
Arachnodes bicolor
Arachnodes didiensis
Arachnodes histeroides
Arachnodes apotolamproides
Arachnodes vieui
Arachnodes vicinus
Arachnodes jeanneli
Arachnodes colasi
Arachnodes refulgens
Arachnodes andriai
Arachnodes sakarahae
Arachnodes splendidus
Arachnodes abadiei
Arachnodes semichalceus
Arachnodes globulosus
Arachnodes renaudi
Arachnodes sicardi
Arachnodes delphinensis
Arachnodes kelifelyi
Arachnodes zombitsyensis
Arachnodes antoetrae
Arachnodes fulgens
Arachnodes neonitidus
Arachnodes philippi
Arachnodes pusillus
Arachnodes tibialis
Arachnodes micheli
Arachnodes prasinus
Arachnodes oberthuri
Arachnodes viettei
Arachnodes micans
Arachnodes genieri
Arachnodes mahafalyensis
Arachnodes dichrous
Arachnodes robinsoni
Arachnodes aeneoviridis
Arachnodes emmae
Arachnodes pillula
Arachnodes dieganus
Arachnodes purpuricollis
Arachnodes mantasoae
Arachnodes balianus
Arachnodes saprinoides
Arachnodes insularis
Arachnodes mantillerii
Arachnodes lendidus
Arachnodes hanskii
Arachnodes seminitidus
Arachnodes ampasindavae
Arachnodes cuprarius
Arachnodes grossepunctatus
Arachnodes vigilans
Arachnodes delaunayi
Arachnodes nitidus
Arachnodes morati
Arachnodes semipunctatus
Arachnodes nosybeensis
Arachnodes luctuosus
Arachnodes manomboensis
Arachnodes seyrigi
Arachnodes manaitrai
Arachnodes ruteri
Arachnodes variolosus
Arachnodes soganus
Arachnodes rubrolimbatus
Arachnodes biimpressus
Arachnodes brunnipes
Arachnodes globuloides
Arachnodes rubrotinctus
Arachnodes morio
Arachnodes pierrettae
Pycnopanelus
→
(3 species)
Pycnopanelus krikkeni
Pycnopanelus rotundus
Pycnopanelus parvicollis
Cambefortantus
→
(8 species)
Cambefortantus micros
Cambefortantus ranomandryi
Cambefortantus ankaratrae
Cambefortantus myops
Cambefortantus helenae
Cambefortantus blanci
Cambefortantus ranomafanensis
Cambefortantus pauliani
Apotolamprus
→
(32 species)
Apotolamprus quadrinotatus
Apotolamprus orangeaensis
Apotolamprus helenae
Apotolamprus bemarahaensis
Apotolamprus marojejyensis
Apotolamprus milloti
Apotolamprus pseudoquadrimaculatus
Apotolamprus darainaensis
Apotolamprus perinetensis
Apotolamprus rubromaculatus
Apotolamprus sericeus
Apotolamprus mahajangaensis
Apotolamprus hanskii
Apotolamprus vadoni
Apotolamprus sahatezaensis
Apotolamprus manjarivoloensis
Apotolamprus sinuatotibialis
Apotolamprus antsingyensis
Apotolamprus cyanescens
Apotolamprus latipennis
Apotolamprus sparsepunctatus
Apotolamprus nanoides
Apotolamprus metallicus
Apotolamprus andranobeensis
Apotolamprus manomboensis
Apotolamprus resplendens
Apotolamprus pseudomanomboensis
Apotolamprus densepunctatus
Apotolamprus quadrimaculatus
Apotolamprus ambohitsitondronensis
Apotolamprus hagai
Apotolamprus zombitsyensis
Outenikwanus tomentosus
Megathoposoma candezei
Aliuscanthoniola similaris
Endroedyolus paradoxus
Dwesasilvasedis medinae
Saphobiamorpha maorianus
Silvaphilus oubosiensis
Eudinopus dytiscoides
Circellium bacchus
Hammondantus psammophilus
Eurysternini
→
(55 species)
Eurysternus calligrammus
Eurysternus nanus
Eurysternus truncus
Eurysternus marmoreus
Eurysternus rivaldii
Eurysternus sulcifer
Eurysternus caribaeus
Eurysternus navajasi
Eurysternus streblus
Eurysternus gilli
Eurysternus hamaticollis
Eurysternus magnus
Eurysternus cavatus
Eurysternus strigilatus
Eurysternus aeneus
Eurysternus plebejus
Eurysternus cyanescens
Eurysternus ventricosus
Eurysternus obliteratus
Eurysternus balachowskyi
Eurysternus impressicollis
Eurysternus atrosericus
Eurysternus wittmerorum
Eurysternus inca
Eurysternus uniformis
Eurysternus nigrovirens
Eurysternus harlequin
Eurysternus velutinus
Eurysternus mexicanus
Eurysternus cayennensis
Eurysternus sanbornei
Eurysternus arnaudi
Eurysternus jessopi
Eurysternus maya
Eurysternus francinae
Eurysternus parallelus
Eurysternus vastiorum
Eurysternus cambeforti
Eurysternus deplanatus
Eurysternus olivaceus
Eurysternus atrocericus
Eurysternus cyclops
Eurysternus superbus
Eurysternus squamosus
Eurysternus angustulus
Eurysternus hypocrita
Eurysternus foedus
Eurysternus martinsi
Eurysternus inflexus
Eurysternus lanuginosus
Eurysternus fallaciosus
Eurysternus gracilis
Eurysternus howdeni
Eurysternus hirtellus
Eurysternus contractus
Scarabaeini
→
(175 species)
scarabs (
Scarabaeus
) →
(147 species)
Scarabaeus subgenus Scarabaeus
→
(13 species)
Scarabaeus westwoodi
Scarabaeus deludens
Scarabaeus caffer
Scarabaeus thphon
Scarabaeus pius
Scarabaeus sacer
Scarabaeus cicatricosus
Scarabaeus goryi
Scarabaeus viettei
Scarabaeus satyrus
Scarabaeus zambezianus
Scarabaeus galenus
Scarabaeus radama
Scarabaeolus
→
(47 species)
Scarabaeus flavicornis
Scarabaeus bohemani
Scarabaeus rugosus
Scarabaeus rubripennis
Scarabaeus rusticus
Scarabaeolus namibensis
Scarabaeolus carniphilus
Scarabaeolus anderseni
Scarabaeolus intricatus
Scarabaeolus inoportunus
Scarabaeolus ebenus
Scarabaeolus inquisitus
Scarabaeolus ermienae
Scarabaeolus andreaei
Scarabaeolus canaliculatus
Scarabaeolus scholtzi
Scarabaeolus soutpansbergensis
Scarabaeolus reichei
Scarabaeolus davisianus
Scarabaeolus obsoletepunctatus
Scarabaeolus megaparvulus
Scarabaeolus lizleri
Scarabaeolus afronitidus
Scarabaeolus kwiluensis
Scarabaeolus parvulus
Scarabaeolus lucidulus
Scarabaeolus cunene
Scarabaeolus karrooensis
Scarabaeolus similis
Scarabaeolus interstitialis
Scarabaeolus knobeli
Scarabaeolus pabulator
Scarabaeolus gracai
Scarabaeolus fritschi
Scarabaeolus clanceyi
Scarabaeolus rugosipennis
Scarabaeolus damarensis
Scarabaeolus funebris
Scarabaeolus laevifrons
Scarabaeolus bohemani
Scarabaeolus krugeri
Scarabaeolus niemandi
Scarabaeolus fragilis
Scarabaeolus werneri
Scarabaeolus orientalis
Scarabaeolus gilleti
Scarabaeolus planipennis
Ateuchetus
→
(9 species)
Scarabaeus laticollis
Ateuchetus savignyi
Ateuchetus semipunctatus
Ateuchetus variolosus
Ateuchetus puncticollis
Ateuchetus armeniacus
Ateuchetus opacipennis
Ateuchetus hottentorum
Ateuchetus catenatus
Sceliages
→
(7 species)
Scarabaeus brittoni
Scarabaeus adamastor
Scarabaeus hippias
Sceliages gagates
Sceliages granulatus
Sceliages difficilis
Sceliages augias
Scarabaeus aranaeoides
Scarabaeus gangeticus
Scarabaeus aegyptiacus
Scarabaeus soutpansbergensis
Scarabaeus winkleri
Scarabaeus nitidicollis
Scarabaeus ambiguus
Scarabaeus rufus
Scarabaeus typhon
Scarabaeus spretus
Scarabaeus bornemisszai
Scarabaeus bellicosus
Scarabaeus megaparvulus
Scarabaeus plansibilis
Scarabaeus licitus
Scarabaeus alienus
Scarabaeus sevoistra
Scarabaeus transcaspicus
Scarabaeus punctatus
Scarabaeus carinatus
Scarabaeus viator
Scarabaeus zambesianus
Scarabaeus zagrosensis
Scarabaeus krutai
Scarabaeus confusus
Scarabaeus fraterculus
Scarabaeus arenarius
Scarabaeus irakensis
Scarabaeus karae
Scarabaeus ascanius
Scarabaeus heqvisti
Scarabaeus carniphilus
Scarabaeus suri
Scarabaeus lanius
Scarabaeus ruficornis
Scarabaeus schulzeae
Scarabaeus basuto
Scarabaeus poggei
Scarabaeus rixosus
Scarabaeus pan
Scarabaeus reichei
Scarabaeus convexus
Scarabaeus fasciatus
Scarabaeus karlwerneri
Scarabaeus acuticollis
Scarabaeus planipennis
Scarabaeus dioscoridis
Scarabaeus jalof
Scarabaeus sculptilis
Scarabaeus inguisitus
Scarabaeus plausibilis
Scarabaeus quadridens
Scarabaeus wilsoni
Scarabaeus tonckeri
Scarabaeus vethi
Scarabaeus isidis
Scarabaeus niemandi
Scarabaeus planifrons
Scarabaeus sulcipennis
Scarabaeus piliventris
Scarabaeus vicinus
Scarabaeus ermienae
Scarabaeus babori
Scarabaeus janssensi
Scarabaeus pabulator
Scarabaeus profanus
Scarabaeus comatus
Scarabaeus cancer
Scarabaeus cognatus
Scarabaeus retusus
Scarabaeus geminogalenus
Drepanopodus
→
(2 species)
Drepanopodus proximus
Drepanopodus costatus
Kheper
→
(25 species)
Kheper kalaharicus
Kheper erichsoni
Kheper sanctus
Kheper bonellii
Kheper venerabilis
Kheper aegyptiorum
Kheper intermedius
Kheper vanderkelleni
Copper dung beetle (
Kheper nigroaeneus
)
Kheper clericus
Kheper aeratus
Kheper zurstrasseni
Kheper laevistriatus
Kheper devotus
Kheper platynotus
Kheper paganus
Kheper rolciki
Kheper cuvieri
Kheper pustulosus
Kheper asceticus
Kheper subaeneus
Dung beetle (
Kheper lamarcki
)
Kheper prodigiosus
Kheper festivus
Kheper porosus
Pachylomerus femoralis
Onthophagus taurus
Other links
Wikidata
Wikipedia (language: en)
Wikipedia (language: es)
Wikipedia (language: fr)
Wikipedia (language: ru)
Wikipedia (language: it)
Wikipedia (language: pt)
Wikipedia (language: ar)
Wikipedia (language: pl)
Wikipedia (language: nl)
Wikipedia (language: fa)
Wikipedia (language: uk)
Wikipedia (language: vi)
Wikipedia (language: ko)
Encyclopedia of Life
NCBI (genetics)
"
Open Tree of Life