Base of the evolutionary tree for Scarabaeini

← parent

• Scarabaeini (175 species)
  • scarabs (Scarabaeus) → (147 species)
    • Scarabaeus subgenus Scarabaeus → (13 species)
      • Scarabaeus westwoodi
      • Scarabaeus deludens
      • Scarabaeus caffer
      • Scarabaeus thphon
      • Scarabaeus pius
      • Scarabaeus sacer
      • Scarabaeus cicatricosus
      • Scarabaeus goryi
      • Scarabaeus viettei
      • Scarabaeus satyrus
      • Scarabaeus zambezianus
      • Scarabaeus galenus
      • Scarabaeus radama
    • Scarabaeolus → (47 species)
      • Scarabaeus flavicornis
      • Scarabaeus bohemani
      • Scarabaeus rugosus
      • Scarabaeus rubripennis
      • Scarabaeus rusticus
      • Scarabaeolus namibensis
      • Scarabaeolus carniphilus
      • Scarabaeolus anderseni
      • Scarabaeolus intricatus
      • Scarabaeolus inoportunus
      • Scarabaeolus ebenus
      • Scarabaeolus inquisitus
      • Scarabaeolus ermienae
      • Scarabaeolus andreaei
      • Scarabaeolus canaliculatus
      • Scarabaeolus scholtzi
      • Scarabaeolus soutpansbergensis
      • Scarabaeolus reichei
      • Scarabaeolus davisianus
      • Scarabaeolus obsoletepunctatus
      • Scarabaeolus megaparvulus
      • Scarabaeolus lizleri
      • Scarabaeolus afronitidus
      • Scarabaeolus kwiluensis
      • Scarabaeolus parvulus
      • Scarabaeolus lucidulus
      • Scarabaeolus cunene
      • Scarabaeolus karrooensis
      • Scarabaeolus similis
      • Scarabaeolus interstitialis
      • Scarabaeolus knobeli
      • Scarabaeolus pabulator
      • Scarabaeolus gracai
      • Scarabaeolus fritschi
      • Scarabaeolus clanceyi
      • Scarabaeolus rugosipennis
      • Scarabaeolus damarensis
      • Scarabaeolus funebris
      • Scarabaeolus laevifrons
      • Scarabaeolus bohemani
      • Scarabaeolus krugeri
      • Scarabaeolus niemandi
      • Scarabaeolus fragilis
      • Scarabaeolus werneri
      • Scarabaeolus orientalis
      • Scarabaeolus gilleti
      • Scarabaeolus planipennis
    • Ateuchetus → (9 species)
      • Scarabaeus laticollis
      • Ateuchetus savignyi
      • Ateuchetus semipunctatus
      • Ateuchetus variolosus
      • Ateuchetus puncticollis
      • Ateuchetus armeniacus
      • Ateuchetus opacipennis
      • Ateuchetus hottentorum
      • Ateuchetus catenatus
    • Sceliages → (7 species)
      • Scarabaeus brittoni
      • Scarabaeus adamastor
      • Scarabaeus hippias
      • Sceliages gagates
      • Sceliages granulatus
      • Sceliages difficilis
      • Sceliages augias
    • Scarabaeus aranaeoides
    • Scarabaeus gangeticus
    • Scarabaeus aegyptiacus
    • Scarabaeus soutpansbergensis
    • Scarabaeus winkleri
    • Scarabaeus nitidicollis
    • Scarabaeus ambiguus
    • Scarabaeus rufus
    • Scarabaeus typhon
    • Scarabaeus spretus
    • Scarabaeus bornemisszai
    • Scarabaeus bellicosus
    • Scarabaeus megaparvulus
    • Scarabaeus plansibilis
    • Scarabaeus licitus
    • Scarabaeus alienus
    • Scarabaeus sevoistra
    • Scarabaeus transcaspicus
    • Scarabaeus punctatus
    • Scarabaeus carinatus
    • Scarabaeus viator
    • Scarabaeus zambesianus
    • Scarabaeus zagrosensis
    • Scarabaeus krutai
    • Scarabaeus confusus
    • Scarabaeus fraterculus
    • Scarabaeus arenarius
    • Scarabaeus irakensis
    • Scarabaeus karae
    • Scarabaeus ascanius
    • Scarabaeus heqvisti
    • Scarabaeus carniphilus
    • Scarabaeus suri
    • Scarabaeus lanius
    • Scarabaeus ruficornis
    • Scarabaeus schulzeae
    • Scarabaeus basuto
    • Scarabaeus poggei
    • Scarabaeus rixosus
    • Scarabaeus pan
    • Scarabaeus reichei
    • Scarabaeus convexus
    • Scarabaeus fasciatus
    • Scarabaeus karlwerneri
    • Scarabaeus acuticollis
    • Scarabaeus planipennis
    • Scarabaeus dioscoridis
    • Scarabaeus jalof
    • Scarabaeus sculptilis
    • Scarabaeus inguisitus
    • Scarabaeus plausibilis
    • Scarabaeus quadridens
    • Scarabaeus wilsoni
    • Scarabaeus tonckeri
    • Scarabaeus vethi
    • Scarabaeus isidis
    • Scarabaeus niemandi
    • Scarabaeus planifrons
    • Scarabaeus sulcipennis
    • Scarabaeus piliventris
    • Scarabaeus vicinus
    • Scarabaeus ermienae
    • Scarabaeus babori
    • Scarabaeus janssensi
    • Scarabaeus pabulator
    • Scarabaeus profanus
    • Scarabaeus comatus
    • Scarabaeus cancer
    • Scarabaeus cognatus
    • Scarabaeus retusus
    • Scarabaeus geminogalenus
  • Drepanopodus → (2 species)
    • Drepanopodus proximus
    • Drepanopodus costatus
  • Kheper → (25 species)
    • Kheper kalaharicus
    • Kheper erichsoni
    • Kheper sanctus
    • Kheper bonellii
    • Kheper venerabilis
    • Kheper aegyptiorum
    • Kheper intermedius
    • Kheper vanderkelleni
    • Copper dung beetle (Kheper nigroaeneus)
    • Kheper clericus
    • Kheper aeratus
    • Kheper zurstrasseni
    • Kheper laevistriatus
    • Kheper devotus
    • Kheper platynotus
    • Kheper paganus
    • Kheper rolciki
    • Kheper cuvieri
    • Kheper pustulosus
    • Kheper asceticus
    • Kheper subaeneus
    • Dung beetle (Kheper lamarcki)
    • Kheper prodigiosus
    • Kheper festivus
    • Kheper porosus
  • Pachylomerus femoralis

Other links

• Wikidata
• Wikipedia (language: en)
• Wikipedia (language: ru)
• Wikipedia (language: pt)
• Wikipedia (language: uk)
• Encyclopedia of Life
• NCBI (genetics)"
• Open Tree of Life

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