OneZoom: Text tree

Base of the evolutionary tree for beetles (Coleoptera)

beetles (Coleoptera) (283406 species)
- Leaf beetles, rove beetles, scarab beetles, longhorn beetles, snout beetles, etc. (Polyphaga) → (237599 species)
  - unnamed group 1642763 → (236888 species)
    - unnamed group 1756750 → (122901 species)
      - unnamed group 1783640 → (96011 species)
      - unnamed group 1756751 → (26890 species)
    - unnamed group 1642764 → (113987 species)
      - unnamed group 1685922 → (70829 species)
      - unnamed group 1642765 → (43158 species)
  - unnamed group 1642054 → (710 species)
    - unnamed group 1642147 → (617 species)
      - unnamed group 1642175 → (589 species)
      - Plate-thigh beetle (Eucinetidae) → (28 species)
    - unnamed group 1642055 → (93 species)
      - Minute beetle (Clambidae) → (57 species)
      - Derodontidae → (36 species)
  - Rhinorhipus tamborinensis
- unnamed group 1596395 → (45658 species)
  - ground beetles (Adephaga) → (45461 species)
    - Caraboidea → (39933 species)
      - Lebiinae → (4454 species)
      - Oxylobus → (29 species)
      - Theprisa → (3 species)
      - Hebeicarabus → (2 species)
      - Licininae → (1354 species)
      - Cryptocephalomorpha → (11 species)
      - Pseudillaphanus → (2 species)
      - Netrodera → (3 species)
      - Caecocaelus → (9 species)
      - Yukihikous → (3 species)
      - Yamautidius → (26 species)
      - Morphodactyla → (6 species)
      - Xystosomus → (9 species)
      - Luzonotrechus → (6 species)
      - Violagonum → (3 species)
      - Loxocrepis → (4 species)
      - Helluodema → (2 species)
      - Eocnides → (2 species)
      - Piezia → (18 species)
      - Platyagonum → (2 species)
      - Rhytidophaena → (4 species)
      - Nothocascellius → (2 species)
      - Eremolestes → (2 species)
      - Sinotrechiama → (6 species)
      - Tapinoscaris → (10 species)
      - Shilinotrechus → (2 species)
      - Archastes → (28 species)
      - Ishidatrechus → (2 species)
      - Calopachys → (4 species)
      - Rhopalomelus → (4 species)
      - Lioptera → (12 species)
      - Pachyroxochus → (3 species)
      - Oxyodontus → (2 species)
      - Leptosarcus → (2 species)
      - Mochtheroides → (4 species)
      - Neonomius → (3 species)
      - Ledouxius → (7 species)
      - Allocota → (4 species)
      - Plagiopyga → (13 species)
      - Bascanus → (10 species)
      - Herculagonum → (3 species)
      - Procletus → (13 species)
      - Sinaphaenops → (13 species)
      - Stygiotrechus → (20 species)
      - Platyxythrius → (20 species)
      - Shiqianaphaenops → (2 species)
      - Speluncarius → (26 species)
      - Lioholus → (2 species)
      - Ardistomis → (49 species)
      - Gigadema → (13 species)
      - Cinctagonum → (4 species)
      - Henvelik → (2 species)
      - Coarazuphium → (11 species)
      - Nebrioferonia → (2 species)
      - Libotrechus → (2 species)
      - Percolaus → (2 species)
      - Cyclosomus → (13 species)
      - Holcocoleus → (2 species)
      - Cothresia → (4 species)
      - Dontolobus → (9 species)
      - Euschizomerus → (16 species)
      - Martyr → (2 species)
      - Iason → (7 species)
      - Pephrica → (6 species)
      - Neodhysores → (2 species)
      - Rhytidognathus → (2 species)
      - Alanorites → (4 species)
      - Eccoptomenus → (3 species)
      - Cintaroa → (2 species)
      - Caecidium → (2 species)
      - Dalmatoreicheia → (2 species)
      - Teropha → (2 species)
      - Oosoma → (2 species)
      - Guiodytes → (3 species)
      - Dyscherus → (14 species)
      - Microzargus → (7 species)
      - Xanthos → (2 species)
      - Aristolebia → (15 species)
      - Gnaphon → (3 species)
      - Agonidium → (49 species)
      - Pseudophorticus → (4 species)
      - Thopeutica → (95 species)
      - Ponapagonum → (2 species)
      - Mamesdytes → (2 species)
      - Sphaerotachys → (3 species)
      - Mesus → (5 species)
      - Pseudabarys → (6 species)
      - Rhabidius → (3 species)
      - Cyanotarus → (2 species)
      - Steganomma → (3 species)
      - Coreoblemus → (5 species)
      - Sinocolpodes → (3 species)
      - Agonica → (4 species)
      - Euryarthron → (21 species)
      - Dromidea → (3 species)
      - Laccopterum → (15 species)
      - Camptotoma → (5 species)
      - Pseudabacetus → (2 species)
      - Trachelocyphus → (5 species)
      - Melaenus → (2 species)
      - Pterostichinae → (3191 species)
      - Taenarthrus → (13 species)
      - Sichuanotrechus → (5 species)
      - Dioryche → (18 species)
      - Pseudosphaerodes → (3 species)
      - Psilidius → (2 species)
      - Dystrichothorax → (48 species)
      - Gnathoxys → (17 species)
      - Plicagonum → (3 species)
      - Eunostus → (14 species)
      - Scalidion → (4 species)
      - Dailodontus → (2 species)
      - Protocerapterus → (2 species)
      - Haplaner → (2 species)
      - Geodromus → (2 species)
      - Polpochila → (23 species)
      - Cratogaster → (4 species)
      - Manautea → (4 species)
      - Brachypelus → (14 species)
      - Eustra → (24 species)
      - Abacillius → (2 species)
      - Hiletus → (6 species)
      - Caledyschirius → (5 species)
      - Hikosanoagonum → (5 species)
      - Migadops → (2 species)
      - Pseudorhysopus → (2 species)
      - Chydaeus → (58 species)
      - Omphreus → (19 species)
      - Guineica → (2 species)
      - Caledonomorpha → (10 species)
      - Letouzeya → (2 species)
      - Straneotia → (3 species)
      - Trechicomimus → (2 species)
      - Myrmecodemus → (5 species)
      - Morionidius → (5 species)
      - Plocamostethus → (2 species)
      - Lobocolpodes → (6 species)
      - Neocarenum → (10 species)
      - Eucaerus → (7 species)
      - Masuzoa → (3 species)
      - Tasmanillus → (2 species)
      - Neolaphyra → (4 species)
      - Phloeoxena → (36 species)
      - Zolotarevskyella → (3 species)
      - Striganoviella → (2 species)
      - Geobaenus → (4 species)
      - Prodyscherus → (21 species)
      - Panagrius → (2 species)
      - Siagonyx → (3 species)
      - Porocimmerites → (10 species)
      - Typsiharpalus → (3 species)
      - Chaetodactyla → (26 species)
      - Sumatrechus → (2 species)
      - Erephognathus → (3 species)
      - Ryugadous → (12 species)
      - Microferonia → (10 species)
      - Negreum → (15 species)
      - Stereostoma → (28 species)
      - Andrewesia → (3 species)
      - Rhytiferonia → (8 species)
      - Systolocranius → (28 species)
      - Collagonum → (11 species)
      - Paussopsis → (2 species)
      - Prioniomus → (15 species)
      - Nothanillus → (2 species)
      - Oodinus → (10 species)
      - Aparupa → (15 species)
      - Pseudotaphoxenus → (111 species)
      - Assadecma → (2 species)
      - Micrixys → (2 species)
      - Trichopselaphus → (8 species)
      - Metallica → (6 species)
      - Thliboclivina → (4 species)
      - Lobatodes → (2 species)
      - Jujiroa → (27 species)
      - Lepcha → (14 species)
      - Cainogenion → (12 species)
      - Gastragonum → (10 species)
      - Nickerlea → (4 species)
      - Camptoscelis → (2 species)
      - Acutosternus → (2 species)
      - Lassalleianum → (2 species)
      - Anisostichus → (4 species)
      - Geballusa → (6 species)
      - Anheterus → (3 species)
      - Callistomimus → (81 species)
      - Metaxymorphus → (28 species)
      - Schuelea → (3 species)
      - Smeringocera → (6 species)
      - Helluosoma → (4 species)
      - Harpalomimetes → (5 species)
      - Dromoceryx → (3 species)
      - Horniulus → (2 species)
      - Nyctosyles → (3 species)
      - Coptolobus → (6 species)
      - Hoplolenus → (4 species)
      - Paulianoscirtus → (2 species)
      - Penthophonus → (5 species)
      - Simodontus → (18 species)
      - Orophicus → (2 species)
      - Metaxys → (18 species)
      - Elliptoleus → (11 species)
      - Adotela → (17 species)
      - Cryptobatis → (8 species)
      - Enantiola → (4 species)
      - Zeopoecilus → (3 species)
      - Oxygnathus → (4 species)
      - Liamegalonychus → (8 species)
      - Antarctiola → (4 species)
      - Nipponagonum → (6 species)
      - Liocolpodes → (3 species)
      - Aepus → (4 species)
      - Diabaticus → (4 species)
      - Pachytrechodes → (4 species)
      - Laparhetes → (2 species)
      - Guizhaphaenops → (11 species)
      - Trechiotes → (5 species)
      - Anoncopeucus → (2 species)
      - Chalcochrous → (6 species)
      - Fortagonum → (29 species)
      - Analoma → (4 species)
      - Daiconotrechus → (3 species)
      - Euryscaphus → (7 species)
      - Aptinoderus → (5 species)
      - Carabops → (8 species)
      - Lipostratia → (9 species)
      - Cuneipectus → (2 species)
      - Crepidopterus → (13 species)
      - Tilius → (6 species)
      - Amblygnathus → (25 species)
      - Alpiodytes → (2 species)
      - Menigius → (9 species)
      - Kultianella → (2 species)
      - Lebistina → (14 species)
      - Pronyssa → (7 species)
      - Cratoferonia → (3 species)
      - Glyptogrus → (7 species)
      - Brachygnathus → (7 species)
      - Cratohaerea → (4 species)
      - Oodinkosa → (3 species)
      - Dister → (2 species)
      - Inkosa → (2 species)
      - Philoscaphus → (6 species)
      - Setalis → (3 species)
      - Reicheadella → (7 species)
      - Leleupidia → (10 species)
      - Molopsida → (6 species)
      - Coptoglossus → (5 species)
      - Altagonum → (39 species)
      - Andinorites → (7 species)
      - Pleuracanthus → (4 species)
      - Solenogenys → (3 species)
      - Ogygium → (2 species)
      - Aspidoglossa → (25 species)
      - Stevensius → (8 species)
      - Machozetus → (2 species)
      - Setodyschirius → (13 species)
      - Scolyptus → (5 species)
      - Velindopsis → (3 species)
      - Lobodontus → (12 species)
      - Meropalpus → (3 species)
      - Omophroninae → (71 species)
      - Kusumia → (22 species)
      - Apophylon → (2 species)
      - Derostichus → (2 species)
      - Paraglycia → (6 species)
      - Meganophthalmus → (5 species)
      - Leleuporites → (2 species)
      - Atrotus → (18 species)
      - Pyramoides → (2 species)
      - Whiteheadiana → (4 species)
      - Bleusea → (2 species)
      - Ancus → (5 species)
      - Kupeharpalus → (3 species)
      - Polychaetus → (2 species)
      - Notopseudomorpha → (2 species)
      - Dhanya → (7 species)
      - Wahlbergiana → (4 species)
      - Oesyperus → (3 species)
      - Andrewesius → (26 species)
      - Ochtozetus → (2 species)
      - Troglorites → (2 species)
      - Junnanotrechus → (9 species)
      - Anillopsis → (2 species)
      - Grouvellina → (21 species)
      - Metallanchista → (2 species)
      - Progonochaetus → (51 species)
      - Cylindronotum → (7 species)
      - Suzuka → (3 species)
      - Eurycoleus → (9 species)
      - Eccoptoptera → (2 species)
      - Meonochilus → (6 species)
      - Chlaeminus → (16 species)
      - Caecoparvus → (14 species)
      - Neoblemus → (10 species)
      - Rugiluclivina → (5 species)
      - Procarabus → (5 species)
      - Neoferonia → (9 species)
      - Austroglyptolenus → (2 species)
      - Loxomerus → (4 species)
      - Archagonum → (2 species)
      - Trirammatus → (11 species)
      - Neodipnus → (4 species)
      - Pogonodaptus → (3 species)
      - Discoptera → (5 species)
      - Zapotecanillus → (8 species)
      - Parareicheia → (3 species)
      - Oxydrepanus → (11 species)
      - Potamagonum → (4 species)
      - Chaetomargoreicheia → (3 species)
      - Goniocellus → (3 species)
      - Adischissus → (14 species)
      - Archidela → (4 species)
      - Sphaerodinus → (2 species)
      - Deltomerodes → (13 species)
      - Semiclivina → (8 species)
      - Simous → (10 species)
      - Physolaesthus → (9 species)
      - Parachlaenius → (11 species)
      - Apocimmerites → (2 species)
      - Nesiocolpodes → (2 species)
      - Montagonum → (9 species)
      - Duvalioblemus → (2 species)
      - Macroprotus → (2 species)
      - Madecassina → (12 species)
      - Epigraphus → (7 species)
      - Metadromius → (30 species)
      - Malagasydipnus → (3 species)
      - Cratognathus → (5 species)
      - Abrodiella → (2 species)
      - Cathaiaphaenops → (6 species)
      - Strigia → (2 species)
      - Reflexisphodrus → (11 species)
      - Hormacrus → (2 species)
      - Parabroscus → (2 species)
      - Abacophrastus → (7 species)
      - Scotoplanetes → (2 species)
      - Trechinites → (2 species)
      - Erectocolliuris → (6 species)
      - Glycia → (6 species)
      - Platycoelus → (18 species)
      - Trichillinus → (22 species)
      - Tantillus → (2 species)
      - Novillidius → (3 species)
      - Gallerucidia → (5 species)
      - Disphericus → (18 species)
      - Harpalomorphus → (7 species)
      - Mexisphodrus → (12 species)
      - Halocoryza → (3 species)
      - Ishikawatrechus → (31 species)
      - Cimmerites → (14 species)
      - Brachichila → (7 species)
      - Velindastus → (2 species)
      - Awatrechus → (11 species)
      - Graniger → (2 species)
      - Rhaebolestes → (2 species)
      - Trechinae → (5039 species)
      - Marsyas → (18 species)
      - Euchroa → (38 species)
      - Oroblemus → (7 species)
      - unnamed group 1620419 → (4180 species)
      - Stilboma → (2 species)
      - Masuzonoblemus → (2 species)
      - Paranillopsis → (2 species)
      - Pseudotoglossa → (7 species)
      - Nannotrechus → (11 species)
      - Idacarabus → (4 species)
      - Brzoskaicheila → (2 species)
      - Feroniola → (10 species)
      - Negrea → (11 species)
      - Duvaliopsis → (7 species)
      - Lachnothorax → (7 species)
      - Agelaea → (2 species)
      - Dyscherinus → (3 species)
      - Giachinoana → (2 species)
      - Holconotus → (8 species)
      - Gracilanillus → (6 species)
      - Amblystogenium → (2 species)
      - Cyrtomoscelis → (21 species)
      - Afrotarus → (10 species)
      - Promegalonychus → (11 species)
      - Kareya → (8 species)
      - Onycholabis → (7 species)
      - Mouhotia → (3 species)
      - Pseudobatenus → (3 species)
      - Agatoides → (2 species)
      - Elaphrinae → (56 species)
      - Metabacetus → (8 species)
      - Syleter → (8 species)
      - Xenodochus → (9 species)
      - Langea → (3 species)
      - Dilonchus → (2 species)
      - Ophonichius → (3 species)
      - Baenningeria → (2 species)
      - Argutoridius → (9 species)
      - Selenotichnus → (3 species)
      - Batoscelis → (12 species)
      - Eurypercus → (3 species)
      - Dactylotrechus → (3 species)
      - Brachynillus → (3 species)
      - Vachinius → (12 species)
      - Haplobothynus → (2 species)
      - Cribrodyschirius → (8 species)
      - Singiliomimus → (2 species)
      - Deipyrodes → (2 species)
      - Nemotarsus → (9 species)
      - Inotrechus → (2 species)
      - Amphimenes → (17 species)
      - Epigraphodes → (3 species)
      - Paropisthius → (4 species)
      - Stichonotus → (4 species)
      - Merizomena → (10 species)
      - Actenoncus → (3 species)
      - Eobroscus → (4 species)
      - Coptia → (4 species)
      - Metagonum → (3 species)
      - Carabilarva → (2 species)
      - Maoritrechus → (3 species)
      - Habutarus → (23 species)
      - Geoffreyella → (2 species)
      - Obadius → (2 species)
      - Pseudoclivina → (12 species)
      - Cryptoscaphus → (2 species)
      - Catacolpodes → (30 species)
      - Allocinopus → (7 species)
      - Leptocolpodes → (12 species)
      - Actodus → (2 species)
      - Forcipator → (4 species)
      - Orionella → (3 species)
      - Climax → (3 species)
      - Accoella → (2 species)
      - Zeanillus → (3 species)
      - Paragonotrechus → (4 species)
      - Miltodes → (2 species)
      - Paramegalonychus → (11 species)
      - Allophanes → (4 species)
      - Helluonidius → (8 species)
      - Protopidius → (5 species)
      - Onypterygia → (34 species)
      - Trilophus → (28 species)
      - Neoglyptus → (7 species)
      - Skorlagad → (4 species)
      - Kupetrechus → (3 species)
      - Gynandromorphus → (2 species)
      - Idiomelas → (4 species)
      - Helluodes → (3 species)
      - Typhlozuphium → (2 species)
      - Feroniascus → (4 species)
      - Teraphis → (6 species)
      - Pilades → (4 species)
      - Serratotyphlus → (2 species)
      - Semiardistomis → (21 species)
      - Strigomerodes → (7 species)
      - Helluopapua → (2 species)
      - Brithysternum → (3 species)
      - Polydamasium → (2 species)
      - Balligratus → (4 species)
      - Crepidonellus → (5 species)
      - Lucicolpodes → (19 species)
      - Pediomorphus → (5 species)
      - Cyphocoleus → (22 species)
      - Eujalmenus → (4 species)
      - Microderes → (10 species)
      - Oreodicastes → (7 species)
      - Haptoderodes → (2 species)
      - Euprosopus → (2 species)
      - Sparostes → (2 species)
      - Incatrechus → (4 species)
      - Creagris → (9 species)
      - Zhijinaphaenops → (5 species)
      - Lebidia → (5 species)
      - Bohemaniella → (4 species)
      - Hartonymus → (2 species)
      - Stolonis → (19 species)
      - Kurasawatrechus → (36 species)
      - Ronhuberia → (2 species)
      - Eucamaragnathus → (15 species)
      - Euryaptus → (6 species)
      - Mnuphorus → (11 species)
      - Hesperanillus → (3 species)
      - Eurylychnus → (8 species)
      - Dimorphoreicheia → (3 species)
      - Chaudoirina → (3 species)
      - Deltomerus → (75 species)
      - Idiastes → (2 species)
      - Sumlinia → (2 species)
      - Nebriinae → (743 species)
      - Ectinothorax → (6 species)
      - Notospeophonus → (3 species)
      - Pseudomonnea → (3 species)
      - Siopelophonus → (2 species)
      - Crenulostrigus → (2 species)
      - Meladroma → (4 species)
      - Mamboicus → (11 species)
      - Dendragonum → (6 species)
      - Teratotarsa → (4 species)
      - Minuthodes → (21 species)
      - Helluocolpodes → (4 species)
      - Setolebia → (5 species)
      - Orthocerodus → (5 species)
      - Paraphaea → (4 species)
      - Anomostomus → (3 species)
      - Rhytisternus → (24 species)
      - Chaetauchenium → (3 species)
      - Pachycarus → (7 species)
      - Pseudagonica → (13 species)
      - Diceromerus → (2 species)
      - Eumara → (4 species)
      - Uenotrechus → (5 species)
      - Amorphomerus → (2 species)
      - Iridagonum → (9 species)
      - Haplopeza → (3 species)
      - Austrotrechus → (2 species)
      - Uenoites → (6 species)
      - Xestagonum → (135 species)
      - Melanchiton → (53 species)
      - Derocrania → (16 species)
      - Catadromus → (8 species)
      - Madascaris → (5 species)
      - Migadopidiella → (2 species)
      - Barylaus → (2 species)
      - Microdaccus → (8 species)
      - Diacanthostylus → (10 species)
      - Tibioscarites → (2 species)
      - Cycloloba → (2 species)
      - Luperca → (2 species)
      - Scaritinae → (1483 species)
      - Strigomerus → (17 species)
      - Otoglossa → (3 species)
      - Harpalinae → (6143 species)
      - Salcedia → (9 species)
      - Camptogenys → (3 species)
      - Luoxiaotrechus → (2 species)
      - Tarastethus → (6 species)
      - Gunvorita → (21 species)
      - Pseudohyparpalus → (20 species)
      - Bothrocolpodes → (4 species)
      - Tropidocerus → (5 species)
      - Trechiama → (176 species)
      - Leleuporella → (5 species)
      - Amblycoleus → (4 species)
      - Eudalia → (13 species)
      - Microodes → (6 species)
      - Anillidius → (7 species)
      - Propionycha → (2 species)
      - Sphoerodes → (4 species)
      - Demetriola → (2 species)
      - Pholeodytes → (5 species)
      - Anchista → (4 species)
      - Kuceraianum → (2 species)
      - Coryza → (10 species)
      - Sloanella → (5 species)
      - Lachnaces → (4 species)
      - Stomonaxellus → (2 species)
      - Straneoa → (2 species)
      - Singilis → (89 species)
      - Neobatenus → (17 species)
      - Ecnomolaus → (2 species)
      - Straneostichus → (7 species)
      - Kaveinga → (23 species)
      - Vianasia → (4 species)
      - Microcephalus → (15 species)
      - Tasmanotrechus → (11 species)
      - Stricklandiana → (11 species)
      - Agastus → (12 species)
      - Chaetobroscus → (3 species)
      - Thoasia → (2 species)
      - Plesioglymmius → (9 species)
      - Hyperectenus → (2 species)
      - Nebrorabus → (3 species)
      - Cheilagona → (6 species)
      - Cerabilia → (11 species)
      - Miscelus → (5 species)
      - Melaeninae → (20 species)
      - Rupa → (5 species)
      - Hyphaereon → (28 species)
      - Valeriaaschero → (2 species)
      - Paussomorphus → (3 species)
      - Cephalostichus → (2 species)
      - Platytarus → (12 species)
      - Dilatotarsa → (8 species)
      - Prosphodrus → (2 species)
      - Moriodema → (2 species)
      - Dalatagonum → (9 species)
      - Crepidolomus → (2 species)
      - Sofota → (5 species)
      - Labocephalus → (4 species)
      - Dicrodontus → (3 species)
      - Bronislavia → (3 species)
      - Paraleleupidia → (27 species)
      - Epikastea → (6 species)
      - Liodaptus → (2 species)
      - Diastrophella → (2 species)
      - Brachyodes → (46 species)
      - Baripus → (25 species)
      - Klapperichella → (2 species)
      - Stylulus → (3 species)
      - Cyrtolaus → (12 species)
      - Megalebia → (2 species)
      - Dromistomus → (5 species)
      - Scototrechus → (3 species)
      - Basilewskya → (3 species)
      - Acheroniotes → (2 species)
      - Parathlibops → (13 species)
      - Chinapenetretus → (9 species)
      - Melanospilus → (5 species)
      - Cymenopterus → (2 species)
      - Cimmeritodes → (3 species)
      - Astastus → (2 species)
      - Matabele → (2 species)
      - Helluomorpha → (7 species)
      - Dicaelindus → (11 species)
      - Parabaris → (3 species)
      - Apsidocnemus → (3 species)
      - Paranchodemus → (4 species)
      - Dyscaris → (4 species)
      - Pseudomegalonychus → (3 species)
      - Metoncidus → (3 species)
      - Kozlovites → (6 species)
      - Cascellius → (2 species)
      - Skouedhirraad → (2 species)
      - Minosaphaenops → (2 species)
      - Xenodromius → (6 species)
      - Colpostoma → (7 species)
      - Abacomorphus → (2 species)
      - Dryptinae → (4 species)
      - Omphreoides → (6 species)
      - Sphodrosomus → (3 species)
      - Cerotalis → (7 species)
      - Arrowina → (8 species)
      - Canarobius → (2 species)
      - Dirotus → (5 species)
      - Epelyx → (5 species)
      - Leptognatha → (33 species)
      - Cynthidia → (6 species)
      - Selenophori → (3 species)
      - Ophoniscus → (8 species)
      - Celioinkosa → (3 species)
      - Agonocheila → (28 species)
      - Thlibops → (2 species)
      - Mimocolliuris → (17 species)
      - Psecadius → (4 species)
      - Queinnectrechus → (14 species)
      - Holcaspis → (35 species)
      - Cosmodiscus → (8 species)
      - Lophocoryza → (3 species)
      - Anisocnemus → (2 species)
      - Yamatosa → (15 species)
      - Trigonothops → (12 species)
      - Metablus → (2 species)
      - Rhyzodiastes → (49 species)
      - Trichopsida → (18 species)
      - Guiaphaenops → (2 species)
      - Gestroania → (4 species)
      - water soldiers (Stratiotes) → (3 species)
      - Shyrodes → (2 species)
      - Asioreicheia → (5 species)
      - Polyaulacus → (4 species)
      - Peronomerus → (6 species)
      - Nycteis → (15 species)
      - Broscini → (220 species)
      - Neochila → (13 species)
      - Allotriopus → (10 species)
      - Satotrechus → (2 species)
      - Moriomorpha → (6 species)
      - Epicolpodes → (6 species)
      - Lorostema → (6 species)
      - Carenidium → (22 species)
      - Paramecus → (4 species)
      - Monocentrum → (11 species)
      - Neodendragonum → (2 species)
      - Sfitakantha → (2 species)
      - Dolichoctis → (125 species)
      - Ooidius → (6 species)
      - Euleptus → (12 species)
      - Eucolliuris → (32 species)
      - Scapterus → (5 species)
      - Naviauxella → (16 species)
      - Trymosternus → (13 species)
      - Laevagonum → (10 species)
      - Abacillodes → (2 species)
      - Maculagonum → (13 species)
      - Macranillus → (5 species)
      - Tarsagonum → (2 species)
      - Mecynoscaris → (2 species)
      - Setalidius → (2 species)
      - Orictites → (2 species)
      - Praepristus → (13 species)
      - Melanchrous → (6 species)
      - Trichocarenum → (3 species)
      - Amphoxyne → (3 species)
      - Hadrocephalus → (3 species)
      - Perissostomus → (2 species)
      - Qianotrechus → (5 species)
      - Holosoma → (10 species)
      - Colasidia → (42 species)
      - Oedesis → (7 species)
      - Myriochile → (3 species)
      - Nanodiodes → (5 species)
      - Sphodropsis → (10 species)
      - Pachycallida → (3 species)
      - Abacetodes → (4 species)
      - Somoplatus → (15 species)
      - Harpalinus → (3 species)
      - Leptagonum → (10 species)
      - Trichocoptodera → (2 species)
      - Helenaea → (3 species)
      - Duvaliomimus → (13 species)
      - Porocara → (5 species)
      - Aepiblemus → (2 species)
      - Neocolpodes → (152 species)
      - Triaenogenius → (11 species)
      - Vata → (2 species)
      - Omobrus → (3 species)
      - Trichisia → (9 species)
      - Oxycentrus → (69 species)
      - Glyptus → (2 species)
      - Naxipenetretus → (2 species)
      - Pelodiaetus → (2 species)
      - Bembidiomorphum → (2 species)
      - Psydrinae → (486 species)
      - Storthodontus → (11 species)
      - Darodilia → (10 species)
      - Skoeda → (11 species)
      - Tostkar → (5 species)
      - Hylotorus → (6 species)
      - Carbonellia → (3 species)
      - Selenodipnus → (3 species)
      - Lonchosternus → (23 species)
      - Zargus → (6 species)
      - Arhytinus → (51 species)
      - Atrirabus → (2 species)
      - Notoplatynus → (2 species)
      - Rakantrechus → (27 species)
      - Kettlotrechus → (5 species)
      - Boeomimetes → (5 species)
      - Axinidium → (11 species)
      - Pterocyrtus → (8 species)
      - Paracoryza → (6 species)
      - Giraffaphaenops → (2 species)
      - Gonogenia → (7 species)
      - Pogonoschema → (4 species)
      - Nesiotrechus → (2 species)
      - Pioprosopus → (5 species)
      - Andrewesinulus → (5 species)
      - Deltocolpodes → (15 species)
      - Delinius → (9 species)
      - Parapatrobus → (2 species)
      - Toshiaphaenops → (2 species)
      - Bedeliolus → (3 species)
      - Nebriagonum → (11 species)
      - Taphoxenus → (10 species)
      - Scaraphites → (7 species)
      - Calyptogonia → (2 species)
      - Puertrechus → (2 species)
      - Meleagros → (4 species)
      - Oribazus → (2 species)
      - Dactyleurys → (3 species)
      - Mexaphaenops → (8 species)
      - Antilliscaris → (4 species)
      - Mizotrichus → (10 species)
      - Thryptocerus → (5 species)
      - Trichis → (3 species)
      - Scaurotrechodes → (3 species)
      - Siagoninae → (83 species)
      - Iresia → (13 species)
      - Aspidaphaenops → (5 species)
      - Bruskespar → (3 species)
      - Camptodontus → (14 species)
      - Parasiopelus → (2 species)
      - Trachysarus → (10 species)
      - Winklerites → (22 species)
      - Orthotrichus → (98 species)
      - Notocolpodes → (33 species)
      - Dendromigadops → (2 species)
      - Eutrechus → (4 species)
      - Morimotoidius → (5 species)
      - Holcoderus → (26 species)
      - Loricerinae → (18 species)
      - Bryanites → (3 species)
      - Atractonotus → (2 species)
      - Lelis → (5 species)
      - Phloeocarabus → (8 species)
      - Promecognathinae → (3 species)
      - Protrechiama → (4 species)
      - Bhutanillus → (2 species)
      - Zuphioides → (23 species)
      - Parabradycellus → (3 species)
      - Tachysbembix → (2 species)
      - Hakaharpalus → (5 species)
      - Helluo → (2 species)
      - Andinocolpodes → (3 species)
      - Trechosia → (12 species)
      - Eremosphodrus → (2 species)
      - Paradyscherus → (3 species)
      - Pseudaphaenops → (2 species)
      - unnamed group 1605233 → (1674 species)
      - Mesolestes → (45 species)
      - Aristopus → (25 species)
      - Neocoptodera → (13 species)
      - Lachnolebia → (2 species)
      - Putzeysius → (2 species)
      - Thyreopterus → (31 species)
      - Anomophaenus → (8 species)
      - Psammodromius → (4 species)
      - Rathymus → (3 species)
      - Mimovelindopsis → (2 species)
      - Troglocimmerites → (10 species)
      - Oodinotrechus → (3 species)
      - Agonorites → (11 species)
      - Chilotomus → (8 species)
      - Stenolepta → (3 species)
      - Typhloscaris → (22 species)
      - Ushijimaella → (7 species)
      - Aztecarpalus → (9 species)
      - Microcheila → (2 species)
      - Hemitelestus → (2 species)
      - Metaxinidium → (4 species)
      - Goedetrechus → (7 species)
      - Qianaphaenops → (6 species)
      - Blennidus → (129 species)
      - Pionycha → (3 species)
      - Archicolliuris → (29 species)
      - Cophosomorpha → (41 species)
      - Microlestodes → (16 species)
      - Cratocerus → (8 species)
      - Lebidema → (3 species)
      - Microdipnites → (6 species)
      - Lobobrachus → (2 species)
      - Dabatrechus → (2 species)
      - Formosiella → (4 species)
      - Probstia → (3 species)
      - Pseudoplatiella → (13 species)
      - Pareuryaptus → (17 species)
      - Cylindropectus → (2 species)
      - Lovricia → (2 species)
      - Trephionus → (14 species)
      - Caucasorites → (3 species)
      - Pelodiaetodes → (5 species)
      - Minypatrobus → (4 species)
      - Dyschiridium → (4 species)
      - Paranurus → (3 species)
      - Orientoreicheia → (3 species)
      - Ardistomopsis → (5 species)
      - Costitachys → (2 species)
      - Atranopsis → (2 species)
      - Afrosyleter → (6 species)
      - Stricteripus → (4 species)
      - Migadopinae → (15 species)
      - Shanwangicarabus → (3 species)
      - Microzaena → (4 species)
      - Euplatia → (2 species)
      - Ripogenites → (2 species)
      - Sphodroschema → (4 species)
      - Reicheiodes → (26 species)
      - Diamella → (4 species)
      - Sinotroglodytes → (2 species)
      - Idiagonum → (16 species)
      - Dinoscaris → (7 species)
      - Selenochilus → (8 species)
      - Broscodes → (4 species)
      - Homalonesiota → (2 species)
      - Psychristus → (21 species)
      - Abaditicus → (2 species)
      - Helluarchus → (2 species)
      - Apotominae → (32 species)
      - Coptopterina → (7 species)
      - Megamorio → (6 species)
      - Geoscaptus → (5 species)
      - Sphodrus → (2 species)
      - Coptocarpus → (14 species)
      - Dongodytes → (11 species)
      - Haplogaster → (9 species)
      - Onotokiba → (6 species)
      - Allotrechiama → (4 species)
      - Colpoides → (5 species)
      - Oecornis → (2 species)
      - Xenitenus → (16 species)
      - Malagasytyphlus → (4 species)
      - Tuiharpalus → (5 species)
      - Harpaglossus → (4 species)
      - Tecnophilus → (2 species)
      - Pseudaristus → (2 species)
      - Deliaesianum → (6 species)
      - Baeoglossa → (2 species)
      - Kenyacus → (15 species)
      - Neanops → (2 species)
      - Platyninae → (2094 species)
      - Syllectus → (3 species)
      - Anillaspis → (2 species)
      - Parascopodes → (2 species)
      - Atlantomasoreus → (2 species)
      - Helluobrochus → (26 species)
      - Mizotrechus → (18 species)
      - Haplotrachelus → (20 species)
      - Somoplatodes → (2 species)
      - Hexachaetus → (12 species)
      - Haplocolpodes → (9 species)
      - Abacoleptus → (3 species)
      - Paranillus → (13 species)
      - Drymatus tessellatus
      - Zeodera atra
      - Thaioodes piceus
      - Callidadelpha bogotana
      - Palaeoaxinidium orapensis
      - Tianeotrechus trisetosus
      - Wulongoblemus tsuiblemoides
      - Pseudomyas doramasensis
      - Amblops piceus
      - Gulaphaenops leptodiroides
      - Cyptomicrus pollicis
      - Microdipnodes tshuapanus
      - Neoleleupidia kochi
      - Shuangheaphaenops elegans
      - Viridagonum lumawigi
      - Brigalowia setifera
      - Crepidogastrillus curtulus
      - Mecynognathus damelii
      - Tinognathus parviceps
      - Eotrechodes larisae
      - Trichopiezia stenotrachela
      - Playtnus indecentis
      - Platysmodes gambeyi
      - Oarotrechus gracilentus
      - Metaxellus abacetoides
      - Parapionycha lizeri
      - Eulampra miranda
      - Homalomorpha castanea
      - Trechiamiotes siamensis
      - Pachytrachelus cribriceps
      - Chaetosaurus emeishanicola
      - Antarctotrechus balli
      - Shennongotrechus politus
      - Daer ales
      - Hemiplatynus chihuahuae
      - Moirainpa amazona
      - Poecilinus madecassus
      - Jocqueius malawicus
      - Chaetoduvalius saetosus
      - Glyphodactyla femoralis
      - Aristochroodes reginae
      - Xanthomelina zajtzewi
      - Xinbinia foveolata
      - Pangus scaritides
      - Denudirabus exstrius
      - Cintaromorpha dacatrai
      - Andinotrechus naranjoi
      - Pontonoa gounellei
      - Peyrieraselus relictus
      - Minipenetretus quadraticollis
      - Chiapadytes bolivari
      - Typhlonesiotes swaluwenbergi
      - Paraphysodeutera naviauxi
      - Rossjoycea glacialis
      - Australovelinda seriata
      - Neochryopus savagei
      - Aquilex diabolicola
      - Ledouxnebria brisaci
      - Speothalpius grayi
      - Afrotachys brincki
      - Psammocoryza vanemdeni
      - Galicioreicheia struyvei
      - Nepalorthogonius monilicornis
      - Celanida montana
      - Lionedya mongolica
      - Promecoptera marginalis
      - Tangarona pensa
      - Acanthoodes centrosternis
      - Mateuellus troglobioticus
      - Anchotrechus punctipennis
      - Agonotrechiotes longiantennatus
      - Cratosoma pictum
      - Kiwiharpalus townsendi
      - Enceladus gygas
      - Chaetocrepis besckii
      - Neodrypta costigera
      - Microblemus rieae
      - Brachinulus viettei
      - Catalanodytes bellesi
      - Cryptocharidius mandibularis
      - Colposphodrus mirandus
      - Guatemalteca virgen
      - Uenanthracus perigonoides
      - Cicindis horni
      - Rhysocara crassa
      - Neoscaphus simplex
      - Lirabus granulatus
      - Pogonopsis pallida
      - Velinda lirata
      - Neothanes testeus
      - Syletor imerinae
      - Madapelmus elongatus
      - Speagonum mirabile
      - Stigmatachys uvea
      - Holaxinidium fitzsimonsi
      - Galaicodytes caurelensis
      - Petrimagnia horricoma
      - Pseudoselenophorus imitator
      - Sinometrius turnai
      - Cardiomera genei
      - Allophanopsis emarginata
      - Plaumannium denticolle
      - Trogloagonum novaehiberniae
      - Protoscalidion rugiae
      - Camptidius ophthalmicus
      - Guatemalatrechus cuchumatanensis
      - Cryptocolliuris pantosi
      - A Ground Beetle (Tetraleucus picticornis)
      - Feronites velenovskyi
      - Yunnanocarabus litus
      - Licentius mirabilis
      - Evolenes exarata
      - Kosswigia insularis
      - Paregaploa conviva
      - Yanzaphaenops hirundinis
      - Tienmutrechus dispersipunctis
      - Externanillus mcraeae
      - Trichopalpoodes hiekei
      - Afrogehringia endroedyi
      - Stegazopteryx ivimkaensis
      - Bothynoproctus mattoensis
      - Paregraphus krusemani
      - Indiophonus pilosus
      - Pectinitarsus holomelas
      - Calathosoma rubromarginatum
      - Aputrechisibus dubius
      - Leleupanillus uluguruanus
      - Rugitarus puellarum
      - Xenaroswelliana deltaquadrant
      - Nesites mirum
      - Beckeria horni
      - Luyatrechus cuelapensis
      - Torretassoa alfierii
      - Diodercarus arrowi
      - Sophroferonia parvitarsis
      - Allardina katangana
      - Hololeius ceylanicus
      - Arbuckle Cave Ground Beetle (Horologion speokoites)
      - Paraxinidium andreaei
      - Fonsecacarabus placidus
      - Anisodyctylus giganteus
      - Lichnasthenus armiventris
      - Hoplizomenus carinatus
      - Kuschelinus insularis
      - Egadyla antelmei
      - Iyatrechus hisamatsui
      - Neohelluo angulicollis
      - Calathocosmus mirus
      - Rhopalostyla virgata
      - Auchmerus trachys
      - Amerinus linearis
      - Abacaelostus filicornis
      - Sinocimmerites cooteri
      - Orthoglymma wangapeka
      - Tarsitachys bilobus
      - Apterodromites saizi
      - Rhopalochlaenius phalanthus
      - Pseudodiachipteryx expansipennis
      - Metascopus camerunicus
      - Harponixus pubescens
      - Tiferonia parva
      - Apsaustodon segregatus
      - Infernophilus castaneus
      - Neotalus portai
      - Caucasaphaenops molchanovi
      - Platymetopsis overali
      - Clivinopsis strigifrons
      - Henrotiochoromus winkleri
      - Trachelocyphoides setulosus
      - Madagascareicheia elongata
      - Ectomomesa coeca
      - Sinesetosa acugena
      - Lithagonum annulicorne
      - Himalopenetretus franzi
      - Neolovricia ozimeci
      - Morphaeus corosus
      - Leleupaussus tetramerus
      - Wanoblemus wui
      - Calleidomorpha nigroaenea
      - Glenopterus laevigatus
      - Haptoderidius debeckeri
      - Eurycalleida diluta
      - Honduranillus balli
      - Chaetotrechiama procerus
      - Megastylulus pivai
      - Titaresius jeanneli
      - Trichidema capensis
      - Geobius pubescens
      - Basilewskyana villiersi
      - Pelocharis remyi
      - Omestes torta
      - Kupeus arcuatus
      - Amphimasoreus amaroides
      - Leptotrachelon nevermanni
      - Patanitretus pakistanensis
      - Tinoderus singularis
      - Pronyssiformia excoffieri
      - Pseudoplatyderus amblyops
      - Himalaphaenops nishikawai
      - A Ground Beetle (Harpalobrachys leiroides)
      - Thalassobius testaceus
      - Dinodes viridis
      - Paraliagonum longipes
      - Spelaeovulcania canariensis
      - Pterogmus rufipes
      - Miolebidia gaura
      - Agonobembix perrieri
      - Gnopheroides pearsoni
      - Neoorthogonius orientalis
      - Renneria kamouni
      - Himalotrechodes insignis
      - Trichognatha marginipennis
      - Plesioaphaenops annae
      - Falsodromius erythropus
      - Oxygoniola chamaeleon
      - Callidagonum pallidum
      - Dianotrechus gueorguievi
      - Srimara planicollis
      - Vietotrechus minutissimus
      - Tachyxysta howdenorum
      - Agonoriascus lapidicola
      - Proditomus mirus
      - Gourlayia regia
      - Paratrichothorax brevistylis
      - Neoeudalia nigra
      - Waiputrechus cavernicola
      - Gibbozaena mirabilis
      - Derossiella nonveilleri
      - Paussotropus cylindricus
      - Neoodes tazieffi
      - Arame macra
      - Darlingtonica papua
      - Epimicodema mastersii
      - Taiwanotrechus subglobosus
      - Aepsera ferruginea
      - Spelaeodytes mirabilis
      - Phanagnathus overlaeti
      - Pachythecus rubrocupreus
      - Praeanchodemus punctaticeps
      - Carenochyrus titanus
      - Basistichus micans
      - Syrdenoidius spinipes
      - Barrymooreana quadrimaculata
      - Trichamblytelus ovalipennis
      - Colpodichius platyderoides
      - Bafutyphlus tsacasi
      - Cymindoides sculptipennis
      - Cistelagonum kaindi
      - Calophaenoidea arrowi
      - Caelopenetretus crinalis
      - Quasipenetretus berezovskii
      - Cameroniola kryzhanovskiji
      - Oroblemites dromioides
      - Pachylebiodes vadoni
      - Pseudosinurus brunneus
      - Superbotrechus bennetti
      - Catascopellus crassiceps
      - Liochirus cycloderus
      - Dromicoida elegantia
      - Pseudopachylebia relucens
      - Laoblemus crypticus
      - Iga formicina
      - Kaffovatus basilewskyi
      - Grundmannius dispar
      - Notospira phalangioides
      - Laoreicheia bulirschi
      - Crassacantha bidens
      - Velindomimus fasciatus
      - Morimotoides formosus
      - Prakasha amariformis
      - Peripristus ater
      - Physocrotaphus ceylonicus
      - Kaszabellus formosanus
      - Bruskmoal bresk
      - Clivinarchus perlongus
      - Hemiaulax dentipennis
      - Cephalosdrophus marinae
      - Mesocolpodes coptoderoides
      - Lobodontidius congoanus
      - Prodiplous businskyi
      - Dyschiriomimus stackelbergi
      - Mantica horni
      - Doliodactyla janatai
      - Samiriamorpha grace
      - Shuaphaenops parvicollis
      - Platylytron amplipenne
      - Cyrtoderus australasiae
      - Criniventer rufus
      - Albanotrechus beroni
      - Antipionycha puncticollis
      - Ischnagonum carinigerum
      - Notokistus brodiei
      - Liotachys antennatus
      - Magnirabus furvus
      - Platynodes westermanni
      - Tricharnhemia browni
      - Cteatus bruchi
      - Pilbaranillus latibasis
      - Anasis howittii
      - Helluapterus niger
      - Lebina resinata
      - Klepteromimus ornatus
      - Gaioxenus pilipalpis
      - Eopaussus baltians
      - Oxyglychus laeviventris
      - Trechoides fasciatus
      - Carayonites insignis
      - Inpa psydroides
      - Velebitaphaenops giganteus
      - Corsyra fusula
      - Pseudotrechisibus sphaericus
      - Kearophus guyanensis
      - A Ground Beetle (Geopinus incrassatus)
      - Karnes baehri
      - Thyreochaetus hirsutus
      - Pachylebia pallipes
      - Speotarus lucifugus
      - Teiresia umbraculata
      - Trichoxycentrus rugiceps
      - Meileyingia spinosa
      - Notolestus sulcipennis
      - Holoponerus godeffroyi
      - Odontomasoreus humeralis
      - Bathytrechus rueci
      - Angionychus lividus
      - Manumorpha biolat
      - Dichaetochilus bicoloripes
      - Hexaplatarthrus vadoni
      - Pareurydera spinosa
      - Sinis brevicollis
      - Eosphodrus potanini
      - Paratrechiotes ocydromoides
      - Penthus tenebrioides
      - Gugheorites scotti
      - Wanhuaphaenops zhangi
      - Hydrotrechus cantabricus
      - Buderes oberti
      - Angustanillus striatipennis
      - Sunocarabus brunneus
      - Liopasa crepera
      - Pterostillichus caecus
      - Pseudanisotarsus nicki
      - Taurocimmerites dublanskii
      - Rhabdotus reflexus
      - Cedrorum azoricus
      - Nebriosoma fallax
      - Nothodaptus simplex
      - Ametroglossus ater
      - Nesaeoduvalius vailatii
      - Harpostomus opulentus
      - Xenitenopsis jacoti
      - Indocarterus inexspectatus
      - Selina westermanni
      - Elgonotyphlus zoiai
      - Corcyranillus menozzii
      - Speokokosia corneti
      - Notabax monteithi
      - Neohyparpalus parcepunctatus
      - Huoyanodytes tujiaphilus
      - Ectenognathus dryptoides
      - Hippoloetis rufus
      - Physeomorpha vianai
      - Erebotrechus infernus
      - Paracimmerites nanus
      - Enoicus fallax
      - Schidonychus brasiliensis
      - Anomotariella hippocrepis
      - Stuhlmannium mirabile
      - Brachynopterus rufulus
      - Decogmus chalybeus
      - Broter ovicollis
      - Wate longinus
      - Afranillus schuelei
      - Asios vindex
      - Aethocarabus levigatus
      - Feostoma irregulare
      - Afrodipnus transvaalianus
      - Notiokasis chaudoiri
      - Xenion ignitum
      - Cardiaderus chloroticus
      - Cryptorites scotti
      - Harpalophonus italus
      - Ogmophora peringueyi
      - Minimaphaenops lipsae
      - Androzelma gigas
      - Yasunimorpha piranha
      - Foveocrotaphus burmanicus
      - Amelus nigripennis
      - Oechalius ohausi
      - Hyperion schroetteri
      - Paulianolebia rubicunda
      - Alkestis nevermanni
      - Othogonius taiwanicus
      - Stenochoromus montenegrinus
      - Corintascaris ferreirae
      - Triplosarus novaezelandiae
      - Xhosores figuratus
      - Phloeozetus pusillus
      - Mexyclothorax baehri
      - Shenaphaenops humeralis
      - Pseudanchomenus aptinoides
      - Xenocelia mandarina
      - Stenoodes jeanneli
      - Hygranillus kuscheli
      - Macromorphus boisduvali
      - Opisthencentrus dentipennis
      - Escolatrechus bolivianus
      - Thalassoduvalius masidai
      - Rawlinsius papillatus
      - Hannaphota distincta
      - Holoprizus serratus
      - Eurystomis castelnaui
      - Megalopaussus amplipennis
      - Pylartesius strandi
      - Nemaglossa brevis
      - Conexicoxa homora
      - Orthobasis bicolor
      - Wuchangicarabus latus
      - Gidda barthelemyi
      - Apristomimus megacephalus
      - Molopinus pilipes
      - Dregus glebalis
      - Neophygas microcephalus
      - Crassagena depressa
      - Pseudometabletus nevermanni
      - Macrotelus pallipes
      - Obesofemoria chijinqiaoensis
      - Balazucellus hubeicola
      - Takasagoagonum scotus
      - Aplothorax burchellii
      - Hystricosphodrus vailatii
      - Oilea spinalis
      - Bellavalentis kuzugamii
      - Dolichocolpodes mirabilis
      - Stenocnemus mannerheimii
      - Phaenaulax nanus
      - Vulcanophilus calathoides
      - Kenyoreicheia aberdarensis
      - Coptopterella scutellata
      - Rhagadillius aethiopicus
      - Aeolodermus emarginatus
      - Quammenis spectabilis
      - Guyanemorpha spectabilis
      - Eucarterus sparsutus
      - Colfax stevensi
      - Idiocolpodes tsaratanensis
      - Aulacolius triordinatus
      - Mexanillus sbordonii
      - Metazuphium spinangulis
      - Paralovricia beroni
      - Ginema thomasi
      - Harpathaumas priscus
      - Lobodontulus mirei
      - Nunbergites aethiopicus
      - Paracolpodes mauritiensis
      - Nipponaphaenops erraticus
      - Litarthrum browni
      - Kar nepalensis
      - Lachenus impunctipennis
      - Apteroessa grossa
      - Mormolycina oberthueri
      - Maoriharpalus sutherlandi
      - Oxygnathopsis javana
      - Mirachlaenius barbarae
      - Himalosphodrus cnesipus
      - Peruphorticus gulliveri
      - Boreaphaenops angustus
      - Fangshania punctata
      - Nipponosynuchus abnormalis
      - Austranillus macleayi
      - Leoglymmius lignarius
      - Sinocarabus longicornutus
      - Amphimenoides maculatus
      - Mascarenhia subappendiculata
      - Prophenorites marojejyanus
      - Medisores abditus
      - Olegius turkmenicus
      - Reicheidius frondicola
      - Dromiops nanniscus
      - Carabidium dejeanianum
      - Procletodema parallela
      - Mesarthropterus wasmanni
      - Sloanoglymmius planatus
      - Metaxidius brunnipenis
      - Percodermus niger
      - Cicindelites armissanti
      - Anillotarsus tetramerus
      - Ifridytes mateui
      - Askalaphium depressum
      - Lymnaem nigropiceum
      - Habragonum amamioshimense
      - Drymonaxus feanus
      - Temnostega antarctica
      - Bamaroodes cyaneus
      - Indopatrobus bashtai
      - Beronaphaenops paphlagonicus
      - Rhysopus klynstrai
      - Rhysopleura orbicollis
      - Prodyscherodes pauliani
      - Amphithasus truncatus
      - Chaetapatrobus valentinae
      - Myanmadera horii
      - Pachyferonia quadricollis
      - Leuropus rubellus
      - Iberodytes ramiroi
      - Parapenthus solitarius
      - Pontodytes cavazzutii
      - Eurynebria complanata
      - Tremembecarabus rotundus
      - Sbordoniella indagi
      - Perucharidius andinus
      - Taniatrechus setosus
      - Setitachys macrops
      - Beipiaocarabus oblongus
      - Afrodromius indotatus
      - Cerapterites primaevus
      - A Ground Beetle (Gehringia olympica)
      - Platysphyrus tibialis
      - Do holotrichius
      - Gastrogmus ischialis
      - Diplacanthogaster bicolor
      - Polyderidius rapoporti
      - Alachnothorax bruchi
      - Lebistinida pulchra
      - Crepidogastrinus kochi
      - Nothobroscus chilensis
      - Kalchdigor deuvei
      - Pterorhopalus mizotai
      - Sinocalosoma longicornutum
      - Croatotrechus tvrtkovici
      - Thayerella newtoni
      - Paraulacoryssus puertoricensis
      - Oxoides obscurus
      - Diachipteryx paradoxa
      - Paulianites nidicola
      - Achaetocephala atrata
      - Crepidozaena gracilis
      - Abacops rugipennis
      - Pollicobius raymondi
      - Distrigidius apicalis
      - Blenniventer violaceotinctus
      - Platyhelluo weiri
      - Vitagonum apterum
      - Omophagus artus
      - Mayaphaenops sbordonii
      - Yunotrechus diannanensis
      - Tibetopenetretus heinzi
      - Galiciotyphlotes weberi
      - Pawgammm rougemonti
      - Dobodura armata
      - Dinocolpodes emeishanicus
      - Phacocerus piceus
      - Chaetagonum collarti
      - Mimanillus gracilis
      - Tianzhuaphaenops jinshanensis
      - Trephisa parallela
      - Harpalodiodes xanthorhaphus
      - Metagrilium westwoodi
      - Nannoryctes sulcaticeps
      - Tuxtlamorpha tuxtla
      - Bhutanotrechus reflexicollis
      - Columbitrechus subsulcatus
      - Jiulongotrechus pubescens
      - Sidublemus solidus
      - Capabatus raffrayi
      - Protocolliuris coerulans
      - Cyrtopilus madecassus
      - Anillopsidius atlasicus
      - Hypodipnites kochi
      - Onawea pantomelas
      - Pseudamblytelus orbicollis
      - Craspophorus reflexus
      - Casaleius ferrugineus
      - Migadopidius bimaculatus
      - Socotrana labroturrita
      - Xishanocarabus parvus
      - Dercylinus impressus
      - Tonkinaphaenops marinae
      - Pachyodontus languidus
      - Jiangxiaphaenops longiceps
      - Pseudomenarus flavomaculatus
      - Dicropterus brevipennis
      - Protocolpodes tratorius
      - Colpomimus colpodoides
      - Speocolpodes franiai
      - Harapalus viridanus
      - Trogloclivina brehieri
      - Cavicarabus lucensus
      - Phyllodroma luteamaculata
      - Synteratus ovalis
      - Tropicoritus ruwenzorii
      - Haplanister crypticus
      - Secatophus australis
      - Lanxangaphaenops louisi
      - Arthropterites klebsi
      - Zoianillus acutipennis
      - Eriophonus grandiceps
      - Lophidius testaceus
      - Jalzicaphaenops poljaki
      - Italodytes stammeri
      - Tauredon horni
      - Passalidius fortipes
      - Mimozaena virescens
      - Mimophilorhizus chilensis
      - Grandopronotalia carnarvona
      - Ground beetle (Xestonotus lugubris)
      - Haploferonia simplex
      - Rubidiclivina punctatissima
      - Junaphaenops tumidipennis
      - Tonkinoscaris excisicollis
      - Dongodytodes brevipenis
      - Paradolichoctis cordicollis
      - Dalmataphaenops chiarae
      - Stenocheila lacordairei
      - Ochyropus gigas
      - Cryptomma multistriatum
      - Megaloodes politus
      - Allosiopelus punctatipennis
      - Straneoites morogoro
      - Grandipenetretus major
      - Kiwitrechus karenscottae
      - Parorthomus socotranus
      - Neodyschirius cruciatus
      - Eurygnathus latreillei
      - Himalotrechus humeratus
      - Archaeocindis johnbeckeri
      - Rypa uenoi
      - Heteracantha depressa
      - Dactylinius punctipennis
      - Sthenocranion severini
      - Bulirschia janaki
      - Euryabax colossus
      - Somalodromius basilewskyi
      - Chaudoirella reichardti
      - Allendia chilensis
      - Pachycaecus savanicola
      - Monodryxus crassus
      - Colantarctia nitens
      - Speozuphium poultieri
    - unnamed group 1596593 → (5528 species)
      - Dytiscoidea → (4635 species)
      - unnamed group 1596594 → (893 species)
  - unnamed group 1596396 → (197 species)
    - reticulated beetles (Archostemata) → (147 species)
      - Reticulated beetle (Cupedidae) → (139 species)
      - unnamed group 1596447 → (8 species)
    - minute bog beetles (Myxophaga) → (50 species)
      - unnamed group 1596416 → (31 species)
      - Skiff beetle (Hydroscaphidae) → (19 species)
- Aulonocnemidae → (54 species)
  - Aulonocnemis → (42 species)
  - Manjarivolo → (3 species)
  - Ankaratrotrox → (6 species)
  - Palnia → (3 species)
- Eobelidae → (12 species)
  - Arnoldibelus → (6 species)
  - Archaeorrhynchoides → (2 species)
    - Archaeorrhynchoides paradoxopus
    - Archaeorrhynchoides latitarsus
  - Khetoxycorynoides retardatus
  - Microprobelus liuae
  - Longidorsum generale
  - Martynovirhynchus arnoldii
- Eupsilobiidae → (3 species)
  - Eidoreus → (2 species)
    - Eidoreus minutus
    - Eidoreus politus
  - Evolocera championi
- Amarygmiidae → (11 species)
  - Cymatothes → (9 species)
  - Meracantha → (2 species)
    - Meracantha contracta
    - Meracantha lacustris
- Brochocoleidae → (5 species)
  - Pareuryomma → (3 species)
  - Odontomma trachylaena
  - Paraodontomma burmitica
- Aglycyderidae → (2 species)
  - Equiflagrum paklapwanensum
  - Equiflagrum pingchauensum
- Termitotrogidae → (12 species)
  - Termitotrox kenyensis
  - Termitotrox cupido
  - Termitotrox ancoroides
  - Termitotrox consobrinus
  - Termitotrox minutus
  - Termitotrox venus
  - Termitotrox monodi
  - Termitotrox turkanicus
  - Termitotrox maynei
  - Termitotrox usumbaricus
  - Termitotrox vanbruggeni
  - Termitotrox permirus
- Nanophyeidae → (2 species)
  - Palaeonanophyes zherikhini
  - Prionophyes acanthomerus
- Eremazidae → (8 species)
  - Eremazus → (6 species)
  - Yixianscarabaeus → (2 species)
    - Yixianscarabaeus sulcatus
    - Yixianscarabaeus tenuestriatus
- Triboliidae → (7 species)
  - Hypogena canaliculata
  - Hypogena marginalis
  - Hypogena tricornis
  - Hypogena depressa
  - Hypogena marginata
  - Hypogena dejeani
  - Hypogena biimpressa
- Lasiosynidae → (22 species)
  - Lasiosyne → (9 species)
  - Parelateriformius → (4 species)
  - Glaphyropterodes → (2 species)
    - Glaphyropterodes gehreti
    - Glaphyropterodes sibiricus
  - Anacapitis → (4 species)
  - Bupredactyla magna
  - Crassisyne ampla
  - Microsyne lauta
- Boleopsis polinae
- Cryptolarynx vitis
- Crowsoniella relicta
- Tritarsus latus
- Sojanocoleus reticulatus
- Passalopalpus cheni
- Septiventer quadridentatus
- Alloioscarabaeus cheni
- Belohina inexpectata
- Wabbel cerebricavus
- Lebanophytum excellens

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